People

Thomas Quinn, Professor, Aquatic & Fishery Sciences tquinn@uw.edu

Tom Quinn was born in New York City and was interested in natural history and especially fishes from an early age. He kept tropical fish, went fishing, and learned to scuba dive. He graduated from Swarthmore College in Pennsylvania with a B.A. in Biology in 1976, traveled west, and completed his Ph.D. in Fisheries at the University of Washington in 1981. He then spent four years as a post-doctoral fellow at the Canadian Department of Fisheries and Oceans, Pacific Biological Station, in Nanaimo, British Columbia. He joined the faculty in the University of Washington’s School of Fisheries, now School of Aquatic and Fishery Sciences, in 1986, and has been studying the ecology, behavior, evolution, and conservation of salmon and trout, and their ecosystems. He taught many classes, including Scientific Writing and Communication, Fisheries Ecology, and Salmonid Behavior and Life History, and co-taught Aquatic Ecological Research in Alaska with Ray Hilborn and Daniel Schindler. He retired from full-time teaching in 2023 but continues to teach his Salmonid Behavior and Life History class, participate in teaching and field work in Alaska, mentor students, and pursue research projects.

Download Tom’s CV

Please peruse this site for information about Tom’s teaching and research at the University of Washington. For copies of past publications, please go to Publications. View Tom’s current lab members and past lab members below.

  • Roni, P., C. Johnson, T. De Boer, G. Pess, and D. Sear. In press. Interannaul variability in the effects of physical habitat and parentage on Chinook salmon egg-to-fry survival. Canadian Journal of Fisheries and Aquatic Sciences.
  • Johnson, C. L., P. Roni, and G. R. Pess. 2012. Parental effect as a primary factor limiting egg-to-fry survival of spring Chinook salmon in the upper Yakima River Basin. Transactions of the American Fisheries Society 141(5):1295-1309.
  • Johnson, C. L., G. M. Temple, T. N. Pearsons, and T. D. Webster. 2009. An evaluation of data entry error and proofing methods for fisheries data. Transactions of the American Fisheries Society 138(3):593-601.
  • Johnson, C. L., P. Roni, A. Murdoch, M. Hughes, and T. De Boer. In prep. Environmental and parental influences to Spring Chinook incubation survival and development in the Chiwawa River, Washington State.

Former Graduate Students and Post-Docs

  • Anderson, J. H. 2006. Colonization of newly accessible habitat by coho salmon (Oncorhynchus kisutch).
  • Berman, C. H. 1990. Effect of elevated holding temperatures on adult spring chinook salmon reproductive success.
  • Boatright, C. 2003. Timing of migration, spawning, and juvenile emergence by sockeye salmon in Bear Lake, Alaska.
  • Buck, G. 2003. Migratory pathways of maturing sockeye salmon in Bristol Bay, Alaska.
  • Buehrens, T. W. 2011. Growth, movement, survival and spawning habitat of coastal cutthroat trout.
  • Chamberlin, J. 2009. Early marine migratory patterns and the factors that promote resident type behavior of Chinook salmon, Oncorhynchus tshawytscha, in Puget Sound, Washington.
  • Dalton, T. J. 1989. The use of a freshwater trematode as a parasite tag to indicate continental region of origin of ocean-caught steelhead trout.
  • Denton, K. 2008 The utilization of a salmon subsidy by resident char (Salvelinus spp.).
  • Doctor, K. 2008 Fishing out evolution? Spatial and temporal patterns of migration in sockeye salmon
  • Erickson, M. 2005. Vertebrate Communities in Bedrock and Gravel bottomed Streams of the Willapa Basin
  • Havey, M. 2008 Salmon olfaction: Odor detection and imprinting in Oncorhynchus spp. pdf
  • Hendry, A. P. 1995. Sockeye salmon (Oncorhynchus nerka) in Lake Washington: an investigation of ancestral origins, population differentiation, and local adaptation.
  • Hodgson, S. 2000. Marine and freshwater climatic influences on the migratory timing of adult sockeye salmon.
  • Jaecks, T. 2010. Population Dynamics and Trophic Ecology of Dolly Varden in the Iliamna River, Alaska: Life History of Freshwater Fish Relying on Marine Food Subsidies.
  • Johnson, C. 2024. Factors affecting the survival and development of upper Columbia River Chinook salmon.
  • Kinnison, M. T. 1997. Population differentiation in chinook salmon introduced to New Zealand: evidence from morphological, reproductive and early life history characters.
  • Kahler, T. H. 1999. Summer movement and growth of individually marked juvenile salmonids in western Washington streams.
  • Kendall, N. 2007. Long term fishery selection on size and age at maturity in Bristol Bay, Alaska sockeye salmon.
  • Leonetti, F. E. 1996. Habitat attributes, sockeye salmon spawning behavior, and redd site characteristics at island beaches, Iliamna Lake, Alaska.
  • Lincoln, A. E. 2019. Selective consumption of sockeye salmon by brown bears: patterns of partial consumption, scavenging, and implications for fisheries management.
  • Mackey, G. 1999. Investigations of opportunities for genetic and ecological interactions and reproductive success of hatchery and wild steelhead (Oncorhynchus mykiss) in Forks Creek, Washington.
  • Newell, J. 2005. Migration and movement patterns of adult sockeye salmon (Oncorhynchus nerka) in Lake Washington.
  • Nowak, G. 2000. Movement patterns and feeding ecology of cutthroat trout (Oncorhynchus clarki clarki) in Lake Washington.
  • Olson, A. F. 1989. Some aspects of the behavior of chinook salmon, Oncorhynchus tshawytscha, in estuaries.
  • Rhodes, J. S. 1998. Comparative performance of hatchery and naturally reared coho salmon parr in streams: laboratory and field experiments
  • Rich, H. R., Jr. 2006. Effects of climate and density on the distribution, growth, and life history of juvenile sockeye salmon in Iliamna Lake, Alaska.
  • Rohde, J. 2013. Partial migration of Puget Sound Coho Salmon (Oncorhynchus kisutch): Individual and population level patterns.
  • Roni, P. 1992. Life history and spawning habitat in four stocks of large-bodied chinook salmon (Oncorhynchus tshawytscha).
  • Shreffler, D. K. 1989. Temporary residence and foraging by juvenile salmon in a restored estuarine wetland.
  • Thornton, E. 2015. Competition between non-native brook trout and coho salmon in the Elwha River, WA during dam removal.
  • Abrey, C. A. 2005. Variation in the early life history of sockeye salmon (Oncorhynchus nerka): emergence timing, an ontogenetic shift, and population productivity.
  • Anderson, J. 2011. Dispersal and reproductive success of Chinook (Oncorhynchus tshawytscha) and coho (O. kisutch) salmon colonizing newly accessible habitat.
  • Arostegui, M. C. 2019. Nonanadromous life history diversity of rainbow trout (Oncorhynchus mykiss).
  • Austin, C. S. 2020. Salmonid life history, phenology, and distribution within a large river basin.
  • Bond, M. H. 2013. Diversity in migration, habitat use, and growth of Dolly Varden char in Chignik Lakes, Alaska.
  • Carlson, S. M. 2006. Evolutionary effects of bear predation on salmon life history and morphology.
  • Cunningham, C. 2015 (co-supervised with Ray Hilborn). Salmonid selection, evolution, and historical abundance patterns.
  • Dickerson, B. R. 2003. Reproductive success in wild pink salmon, Oncorhynchus gorbuscha.
  • Dittman, A. H. 1994. Behavioral and biochemical mechanisms of olfactory imprinting and homing by coho salmon.
  • Gende, S. M. 2002. Foraging behavior of bears at salmon streams: intake, choice, and the role of salmon life history.
  • Goetz, F.A. 2016. Migration and Residence Patterns of Salmonids in Puget Sound, Washington.
  • Hendry, A. P. 1998. Reproductive energetics of Pacific salmon: strategies, tactics and tradeoffs.
  • Hovel, R. 2015. Species diversity and environmental variability: patterns and processes of
    lacustrine fish community responses in a variable world.
  • Kendall, N. W. 2011. Fishery selection and Pacific salmon life histories: patterns and processes.
  • Kinnison, M. T. 1999. Life history divergence and population structure of New Zealand chinook salmon: a study of contemporary microevolution.
  • Lonzarich, D. G. 1994. Stream fish communities in Washington: patterns and processes.
  • McElroy, K. 2023. Applying the Ideal Free Distribution Theory to two mobile predators on Pacific salmon: Commercial fishers and brown bears.
  • McLean, J. E. 2003.Reproductive success of hatchery and wild steelhead, Oncorhynchus mykiss.
  • Merrick, R. 1995. The relationship of the foraging ecology of Steller sea lions (Eumetopias jubatus) to their population decline in Alaska.
  • Pess, G. R. 2009. Patterns and processes of salmon colonization.
  • Roni, P. 2000. Response of fish and salamanders to instream restoration in small western Washington and Oregon streams.
  • Seamons, T. R. 2005. The mating system of steelhead and the effect of length and arrival date on steelhead reproductive success.
  • Tillotson. M. D. 2018. Impacts of climate change on sockeye salmon life-history productivity and evolutionary ecology: Implications for managing salmon in a warming world.
  • Woods, P. J. 2011. Ecological diversity in the polymorphic fish Arctic charr (Salvelinus alpinus).

Research

My students, post-docs, and I have been studying the behavior, ecology, evolution, and conservation of salmon, trout and char. These fishes are noteworthy for their migrations and homing behavior, complex population structure, importance in commercial, recreational, and subsistence fisheries, roles are cultural icons around the Pacific Rim, and complex connections within their ecosystems. Moreover, some populations are in serious decline or have already become extinct, chiefly towards the southern end of their distribution, although others are healthy and support sustainable fisheries. Therefore, the research that we conduct integrates the natural and human-related processes affecting the fish, and we emphasize the importance of linking basic scientific investigations to conservation and management. The specific projects have varied over the years, as the interests of students, the pressing issues, and the funding opportunities have changed.

Current projects include ones conducted in western Alaska, at the University of Washington’s field camps, and ones conducted in Washington, chiefly in the Puget Sound region. As part of a long-term, multi-investigator program whose goal is to understand the basic and applied ecology of western Alaska sockeye salmon and their habitats [link to ASP website], we have studied the life history traits, spawning site characteristics, reproductive behavior, energetics, and predator-prey interactions of sockeye salmon and other fishes. Specific projects have examined: (1) the scale of population structure and homing by sockeye salmon, (2) the relationships between stream entry date, body size, population density, energetics and longevity of adult sockeye salmon in small streams, (3) the influences of habitat, body size, and sex on vulnerability of adult sockeye salmon to predation by bears, and (4) the forms of selection exerted on salmon by the commercial fishery. In addition to these projects on sockeye salmon, we are also investigating the ecology, migrations, and life history of char in the genus Salvelinus, including Dolly Varden and Arctic char, and rainbow trout.

In Washington, we have been studying the re-colonization of rivers by salmon and trout after dam modification (in the Cedar River) and removal (the Elwha River), including projects investigating estuary, river and lake habitats, in collaboration with scientists representing many different agencies and organizations. We are also investigating the migration patterns of different salmonids species in Puget Sound. This work involves a combination of ultrasonic tagging and tracking, and also analysis of coded wire tagging data, in collaboration with federal and state agency scientists. We are comparing the movement patterns of steelhead trout, cutthroat trout, bull trout, coho salmon and Chinook salmon.

Publications

Austin, C. A., C. E. Torgersen, and T. P. Quinn. Who spawns where? Temperature, elevation, and discharge differentially affect the distribution of breeding by six Pacific salmonids within a large river basin. Canadian Journal of Fisheries and Aquatic Sciences 80: 1365-1384.

Casendino, H. R., K. N. McElroy, M. H. Sorel, T. P. Quinn, and C. L. Wood. 2023. Two decades of change in sea star abundance at a subtidal site in Puget Sound, Washington. PLoS ONE 18(6): e0286384. doi.org/10.1371/journal.pone.0286384

Chalde, T., C. Nardi, J. Reibel, and T. P. Quinn. 2023. Detection of juvenile coho salmon (Oncorhynchus kisutch) indicates widespread successful reproduction and different life-history strategies in the extreme south of Patagonia. Biological Invasions 25:3443–3452.   https://doi.org/10.1007/s10530-023-03116-1

Kendall, N. W., J. Unrein, C. Volk, D. A. Beauchamp, K. L. Fresh, and T. P. Quinn. 2023. Life-cycle model reveals sensitive life stages and evaluates recovery options for a dwindling Pacific salmon population. North American Journal of Fisheries Management 43: 203-230.   

Kiffney, P. M., P. J. Lisi, M. Liermann, S. M. Naman, J. H. Anderson, M. H. Bond, G. R. Pess, M. E. Koehler, E. R. Buhle, T. W. Buehrens, R. S. Klett, J. R. Cram, and T. P. Quinn. 2023. Colonization of a temperate river by mobile fish following habitat reconnection. Ecosphere 2023; e4336 https://doi.org/10.1002/ecs2.4336

Kilfoil, J. P., T. P. Quinn, and A. J. Wirsing. 2023. Human effects on brown bear diel activity may facilitate subadults foraging on Pacific salmon. Global Ecology and Conservation 42: e02407

Minkoff, D., W. R. Ardren, K. Kaiser, A. H. Dittman, T. P. Quinn, J. Atema, and B. W. Taylor. 2023. Spatiotemporal patterns of dissolved free amino acids in New England rivers could be unique and stable odor signatures for imprinting and homing by Atlantic salmon. Freshwater Science 42: 375-391. https://doi.org/10.1086/728120

Pease, J. E., J. P. Losee, S. Caromile, G. Madel, M. Lucero, A. Kagley, M. G. Bertram, J. M. Martin, T. P. Quinn, D. Palm, and G. Hellström. 2023. Comparison of triploid and diploid rainbow trout (Oncorhynchus mykiss) fine-scale movement, migration and catchability in lowland lakes of western Washington. Movement Ecology 11(57): 1-13. https://doi.org/10.1186/s40462-023-00418-w

Rheinsmith, S. E., T. P. Quinn, A. H. Dittman, and K. E. Yopak. 2023. Ontogenetic shifts in olfactory rosette morphology of the sockeye salmon, Oncorhynchus nerka. Journal of Morphology 284(1): e21539. doi.org/10.1002/jmor.21539

Thomas, Z. R., D. A. Beauchamp, C. P. Clark, and T. P. Quinn. 2023. Seasonal shifts in diel vertical migrations by lake-dwelling coastal cutthroat trout, Oncorhynchus clarkii clarkii, reflect thermal regimes and prey distributions. Ecology of Freshwater Fish 32: 842-851. 

Armstrong, M. E., D. Minkoff, A. H. Dittman, D. May, E. K. Moody, T. P. Quinn, J. Atema, and W. R. Ardren.  2022. Evidence of an olfactory imprinting window in embryonic Atlantic salmon. Ecology of Freshwater Fish 31: 270-279.

Chin, A. T., M. H. Bond, and T. P. Quinn. 2022. Life history differences between 2 anadromous populations of the northern form of the Dolly Varden (Salvelinus malma) in Bristol Bay in southwestern Alaska. Fishery Bulletin 120: 234-239.

Dittman, A. H., C. J. Cunningham, and T. P. Quinn. 2022. Can unique amino acid profiles guide adult salmon to natal streams? A comparison of streams sampled prior to and after the arrival of adult Pacific salmon. Hydrobiologia 849: 3501-3513.

Polyakov, A. Y., K. W. Myers. T. P. Quinn, A. M. Berdahl, 2022. Group size affects predation risk and foraging success in Pacific salmon at sea. Science Advances 8:eabm7548.

Quinn, T. P. 222. Book Review: Trout and Salmon of the Genus Salmo. Johannes Schöffmann. American Fisheries Society, Bethesda, Maryland. 2021.

Quinn, T. P. and J. P. Losee. 2022. Diverse and changing use of the Salish Sea by Pacific salmon, trout, and char. Canadian Journal of Fisheries and Aquatic Sciences 79: 1003-1021.

Quinn, T.P., M. D. Scheuerell, J. P. Losee, and D. Hanada. 2022. Multi-decadal trends in body size of Puget Sound Chinook Salmon: Analysis of data from the Tengu Derby, a culturally unique fishery. Marine and Coastal Fisheries 14:e10205 9 pp.

Quinn, T. P., A. J. Wirsing, and M. Proctor. 2022. Optimal barbed wire height for brown bear hair sample collection. Ursus 33: article e2: 1-6.

Waldman, J. R. and T. P. Quinn. 2022. North American diadromous fishes: Drivers of decline and potential for recovery in the Anthropocene. Sciences Advances 8(4): eabl5486 (16 pages).

Waples, R. S, M. J. Ford, K. Nichols, M. Kardos, J. Myers, T. Q. Thompson, E. C. Anderson, I. J. Koch, G. McKinney, M. R. Miller, K. Naish, S.R. Narum, K. G. O'Malley, D. Pearse, G. R. Pess, T. P. Quinn, T. R. Seamons, A. Spidle, K. Warheit, and S. C. Willis. 2022. Implications of large-effect loci for conservation: a review and case study with Pacific salmon. Journal of Heredity 113: 121-144.

Zimmerman, M. S., M. R. Sloat, K. V. Kuzishchin, M. C. Arostegui, M. A. Gruzdeva, T. Seamons, and T. P. Quinn. 2022. Diversity of life history traits, growth, and lipid storage in migratory variants of steelhead and rainbow trout (Oncorhynchus mykiss) in Kamchatka, Russia. Canadian Journal of Fisheries and Aquatic Sciences 79: 1625-1640.

Austin, C. S., T. E. Essington, and T. P. Quinn. 2021. In a warming river, wild Chinook salmon spawn later but hatchery-origin conspecifics do not. Canadian Journal of Fisheries and Aquatic Sciences 78: 68-77.

Goetz, F. A., E. Beamer, E. J. Connor, E. Jeanes, C. Kinsel, J. W. Chamberlin, C. Morello, and T. P. Quinn. 2021. The timing of anadromous bull trout migrations in estuarine and marine waters of Puget Sound, Washington. Environmental Biology of Fishes 104: 1073-1088.

Lincoln, A. E., A. J. Wirsing, and T. P. Quinn. 2021. Prevalence and patterns of scavenging by brown bears on salmon carcasses. Canadian Journal of Zoology 99: 9-17.

Quinn, T. P. 2021. Differential migration in Pacific salmon and trout: Patterns and hypotheses. Animal Migration 8: 1-18.

Quinn, T. P. 2021. Time required for brown bears to capture and consume Pacific salmon. Western North American Naturalist 81: 471-476.

Quinn, T. P., G. R. Pess, B. J. G. Sutherland, S. J. Brenkman, R. E. Withler, K. Flynn, and T. D. Beacham. 2021.. Resumption of anadromy or colonization from afar? Sources of sockeye salmon Oncorhynchus nerka in the Elwha River after dam removal. Transactions of the American Fisheries Society 150: 452-464.

Ro, H., J. H. Stern, A. J. Wirsing, and T. P. Quinn. 2021. Stable isotopes reveal variation in consumption of Pacific salmon by brown bears, despite ready access in small streams. Journal of Fish and Wildlife Management 12: 40-49.

Tillotson, M. D., M. C. Arostegui, C. S. Austin, A. E. Lincoln, W. Matsubu, K. N. McElroy, and T. P. Quinn. 2021. Challenges in the identification and interpretation of phenological shifts: Anthropogenic influences on adult migration timing in salmonids. Reviews in Fisheries Science and Aquaculture 29: 769-790.

Whitmore, M, S. Richardson, A. Huff, K. Goodson, T. P. Quinn, A. H. Dittman, M. A. Johnson, M. Kamran, and D. L. G. Noakes. 2021. Homeward bound: in-river movements of adult hatchery- and natural-origin Chinook Salmon in the Elk River, Oregon. North American Journal of Fisheries Management 41: 1088-1096.

Barnett, H. K., T. P. Quinn, M. Bhuthimethee, and J. R. Winton. 2020. Increased prespawning mortality threatens an integrated natural- and hatchery-origin sockeye salmon population in the Lake Washington Basin. Fisheries Research 227 (105527): 1-10.

Dittman, A. H., and T. P. Quinn. 2020. Amino acid cues emanating from Pacific salmon eggs and ovarian fluid. Journal of Fish Biology 97: 1408-1414.

Levi, T., G. V. Hilderbrand, M. D. Hocking, T. P Quinn, K. S. White, M. S. Adams, J. B. Armstrong, A. P. Crupi, C. T. Darimont, W. Deacy, S. L. Gilbert, W. J. Ripple, Y. Shakeri, R. E. Wheat, C. C. Wilmers. Community ecology and conservation of bear-salmon ecosystems. Frontiers in Ecology and Evolution 8: 513304.

Lincoln, A., R. Hilborn, A. J. Wirsing, and T. P. Quinn. 2020. Managing salmon for wildlife: Do fisheries limit salmon consumption by bears in small streams? Ecological Applications 30(3): e02061

Lincoln, A., A. J. Wirsing, and T. P. Quinn. 2020. Long-term use of noninvasive sampling methods: does successful sampling of brown bears (Ursus arctos) by hair snares change over time? Wildlife Research 47: 499-508.

Michel, C. J., J. M. Smith, B. M. Lehman, N. J. Demetras, D. D. Huff, P. L. Brandes, J. A. Israel, T. P. Quinn, and S. A. Hayes. 2020. Limitations of active removal to manage predatory fish populations. North American Journal of Fisheries Management 40: 3-16.

Taugbøl, A., T. P. Quinn, K. Østbye, L. A. Vøllestad. 2020. Allometric relationships in morphological traits associated with foraging, swimming ability and predator defence reveal adaptations toward brackish and freshwater environments in the threespine stickleback. Ecology and Evolution 10: 13412-13426.

Wirsing, A. J., T. P. Quinn, J. R. Adams, and L. P. Waits. 2020. Optimizing selection of brown bear hair for noninvasive genetic analysis. Wildlife Society Bulletin 44: 94-100.

Wold, K., A. J. Wirsing, and T. P. Quinn. Do brown bears (Ursus arctos) avoid barbed wires deployed to obtain hair samples? A videographic assessment. Wildlife Biology 2020(1):wlb.00664

Arostegui, M. C., and T. P. Quinn. 2019. Ontogenetic and ecotypic variation in the coloration and morphology of rainbow trout (Oncorhynchus mykiss) in a stream-lake system. Biological Journal of the Linnean Society 128: 681-699.

Arostegui, M. C., and T. P. Quinn. 2019. Reliance on lakes by salmon, trout, and charr (Oncorhynchus, Salmo, and Salvelinus): An evaluation of spawning habitats, rearing strategies, and trophic polymorphisms. Fish and Fisheries 20: 775-794

Arostegui, M. C., T. P. Quinn, L. W. Seeb, J. E. Seeb, and G. J. McKinney. 2019. Retention of a chromosomal inversion from an anadromous ancestor provides the genetic basis for alternative freshwater ecotypes in rainbow trout. Molecular Ecology 28:1412–1427

Austin, C. S., M. H. Bond, J. M. Smith, E. D. Lowery, and T. P. Quinn. 2019. Partial migration and life history variation in a facultatively anadromous, iteroparous salmonid, bull trout (Salvelinus confluentus). Environmental Biology of Fishes 102: 95-104

Austin, C. S., T. E. Essington, and T. P. Quinn. 2019. Spawning and emergence phenology of bull trout Salvelinus confluentus under differing thermal regimes. Journal of Fish Biology 94: 191-195.

Axworthy, J. B., J. M. Smith, M. S. Wing, and T. P. Quinn. 2019. Sex biased individual variation in movement patterns of a highly mobile, near-shore marine planktivore, the reef manta ray, Manta alfredi. Journal of Fish Biology 95: 1399-1406.

Barnett, H. K., T. P. Quinn, K. L. Fresh, M. E. Koehler, K. D. Burton, M. Bhuthimethee, N. P. Peterson. 2019. Differential marking of embryos by location and date of release reveals within-river natal homing and parental influence on progeny return timing in Sockeye Salmon. Transactions of the American Fisheries Society 148: 393-405.

Cathcart, C. N., P. A.H. Westley, K. J. Dunker, T. P. Quinn, A. J. Sepulveda, F. A. von Hippel, A. Wizik, and D. B. Young. 2019. Trophic plasticity and the invasion of a renowned piscivore: A diet synthesis of northern pike (Esox lucius) from the native and introduced range of Alaska, U.S.A. Biological Invasions 21: 1379-1392

Feddern, M. L., G. Holtgrieve, J. Hart, S. Perakis, H. Ro, T. P. Quinn. 2019. Riparian soil nitrogen cycling and isotopic enrichment in response to a long-term carcass manipulation experiment. Ecosphere 10(11): e02958.

Goetz, F. A., and T. P. Quinn. 2019. Behavioral thermoregulation by adult Chinook salmon (Oncorhynchus tshawytscha) in estuary and freshwater habitats prior to spawning. Fishery Bulletin 17: 258-271.

Hovel, R. A., K. L. Fresh, S. L. Schroder, A. H. Litt, and T. P. Quinn. 2019. A wide window of migration phenology captures inter-annual variability of favourable conditions: Results from a whole-lake experiment with juvenile Pacific salmon. Freshwater Biology 64: 46-55.

Howard, I., E. Davis, G. Lippert, T. P. Quinn, and C. L. Wood. 2019. Abundance of an economically important nematode parasite increased in Puget Sound between 1930 and 2016: Evidence from museum specimens confirms historical data. Journal of Applied Ecology 56: 190-200.

Lincoln, A. E., and T. P. Quinn. 2019. Optimal foraging or surplus killing: selective consumption and discarding of Pacific salmon by brown bears. Behavioral Ecology 30: 202-212.

Ohlberger, J., S. J. Brenkman, P. Crain, G. R. Pess, J. J. Duda, T. W. Buehrens, T. P. Quinn, and R. Hilborn. 2019. A Bayesian life-cycle model to estimate escapement at MSY in salmon based on limited information. Canadian Journal of Fisheries and Aquatic Sciences 76: 299-307.

Oke, K. B., C. J. Cunningham, T. P. Quinn, and A. P. Hendry. 2019. Independent lineages in a common environment: the roles of determinism and contingency in shaping the migration timing of even- versus odd-year pink salmon over broad spatial and temporal scales. Ecology Letters 22: 1547-1556.

Oke, K. B., E. Motivans, T. P. Quinn, and A. P. Hendry. 2019. Sexual dimorphism modifies habitat-associated divergence: Evidence from beach and creek breeding sockeye salmon. Journal of Evolutionary Biology 32: 227-242.

Sparks, M. M., J. A. Falke, T. P. Quinn, M. D. Adkison, D. E. Schindler, K. Bartz, D. Young, and P. A. H. Westley. 2019. Influences of spawning timing, water temperature, and climatic warming on early life history phenology in western Alaska sockeye salmon. Canadian Journal of Fisheries and Aquatic Sciences 76: 123-135.

Tillotson, M. D., H. K. Barnett, M. Bhuthimethee, M. E. Koehler, and T. P. Quinn. 2019. Artificial selection on reproductive timing in hatchery salmon drives a phenological shift and potential maladaptation to climate change. Evolutionary Applications 12: 1344-1359.

Arostegui, M. C., and T. P. Quinn. 2018. Trophic ecology of nonanadromous rainbow trout in a postglacial lake system: partial convergence of adfluvial and fluvial forms. Canadian Journal of Zoology 96: 818-827.

Arostegui, M.C., R.A. Hovel, and T.P. Quinn. 2018. Schistocephalus solidus parasite prevalence and biomass intensity in threespine stickleback vary by habitat and diet in boreal lakes. Environmental Biology of Fishes. 101: 501-514.

Gomez-Uchida, D., D. Cañas-Rojas, C. Riva-Rossi, J. Ciancio, M. Pascual, B. Ernst, E. Aedo Marchant, S. Musleh, F. Valenzuela-Aguayo, T.P. Quinn, J. Seeb, and L. Seeb. 2018. Genetic signatures of artificial and natural dispersal linked to colonization of South America by non-native Chinook salmon (Oncorhynchus tshawytscha). Ecology and Evolution 8: 6192-6209.

Hasselman, D. J., P. Bentzen, S. Narum, and T. P. Quinn. 2018. Rapid formation of population genetic structure following the introduction and establishment of non-native American shad (Alosa sapidissima) along the Pacific Coast of North America. Biological Invasions 20:3123–3143.

Kiffney, P. M., B. Cram, P. Faulds, K. Burton, M. Koehler, and T. P. Quinn. 2018. Spatiotemporal patterns of mountain whitefish (Prosopium williamsoni) in response to a restoration of longitudinal connectivity. Ecology of Freshwater Fish 27: 1037-1053.

Lincoln, A. E., J. A. Shaffer, and T. P. Quinn. 2018. Opportunistic use of estuarine habitat by juvenile bull trout, Salvelinus confluentus, from the Elwha River before, during, and after dam removal. Environmental Biology of Fishes 101: 1559-1569.

Ohlberger, J., T. W. Buehrens, S. J. Brenkman, P. Crain, T. P. Quinn, and R. Hilborn. 2018. Effects of past and projected river discharge variability on freshwater production in an anadromous fish. Freshwater Biology 63: 331-340.

Putman N.F., M.M. Scanlan, A.M. Pollock, J.P. O’Neil, R.B. Couture, J.S. Stoner, T.P. Quinn, K.J. Lohmann, and D.L.G. Noakes. 2018. Geomagnetic field influences upward movement of young Chinook salmon emerging from nests. Biology Letters.

Quinn, T. P. 2018. The Behavior and Ecology of Pacific Salmon and Trout. University of Washington Press, Seattle. Second edition, 547 pp.

Quinn, T. P. 2018. From magnets to bears: Is a career studying salmon narrow or broad? ICES Journal of Marine Science 75: 1546-1552.

Quinn, T.P., J. M. Helfield, C. Austin, R. Hovel, and A. G. Bunn. 2018. A multi-decade experiment shows that fertilization by salmon carcasses enhanced tree growth in the riparian zone. Ecology 99: 2433-2441.

Tillotson, M.D., R.P. Kelly, J.J. Duda, M. Hoy, J. Kralj, and T.P. Quinn. 2018. Concentrations of environmental DNA (eDNA) reflect spawning salmon abundance at fine spatial and temporal scales. Biological Conservation. 220: 1-11.

Tillotson, M.D., and T.P. Quinn. 2018. Selection on the timing of migration and breeding: A neglected aspect of fishing-induced evolution and trait change. Fish and Fisheries. 19: 170-181.

Wirsing, A. J., T. P. Quinn, C. J. Cunningham, J. R. Adams, A. D. Craig, and L. P. Waits. 2018. Alaskan brown bears (Ursus arctos) aggregate and display fidelity to foraging neighborhoods while preying on Pacific salmon along small streams. Ecology and Evolution 8: 9048-9061.

Woods, P. J., S. Skúlason, S. S. Snorrason, and T. P. Quinn. 2018. Eco-evolutionary feedbacks in the functional role of a polymorphic colonizer: Arctic charr in subarctic lakes of Alaska and Iceland. Evolutionary Ecology Research 19: 149-169.

Arostegui, M.C., J.M. Smith, A.N. Kagley, D. Spilsbury-Pucci, K.L. Fresh, and T.P. Quinn. 2017. Spatially clustered movement patterns and segregation of subadult Chinook salmon within the Salish Sea. Marine and Coastal Fisheries 9(1): 1-12.

Arostegui, M.C., T.E. Essington, and T.P. Quinn. 2017. Interpreting vertical movement behavior with holistic examination of depth distribution: a novel method reveals cryptic diel activity patterns of Chinook salmon in the Salish Sea. Animal Biotelemetry 5:2.

Berdahl, A., P.A.H. Westley, and T.P. Quinn. 2017. Social interactions shape the timing of spawning migrations in an anadromous fish. Animal Behaviour 126: 221-229.

Bond, M.H., P.A. H. Westley, A.H. Dittman, D. Holecek, T. Marsh, and T.P. Quinn. 2017. Combined effects of barge transportation, river environment, and rearing location on straying and migration of adult Snake River fall-run Chinook salmon. Transactions of the American Fisheries Society. 146: 60-73.

Havey, M.A., A.H. Dittman, T.P. Quinn, S.C. Lema, and D. May. 2017. Experimental evidence for olfactory imprinting by sockeye salmon at embryonic and smolt stages. Transactions of the American Fisheries Society 146: 74-83.

Hovel, R.A., S.M. Carlson, and T.P. Quinn. 2017. Climate change alters the reproductive phenology and investment of a lacustrine fish, the three-spine stickleback. Global Change Biology.

Hovel, R.A., J.T. Thorson, J.L. Carter, and T.P. Quinn. 2017. Within-lake habitat heterogeneity mediates community response to warming trends. Ecology 98(9): 2333-2342.

Kagley, A.N., J.M. Smith, K.L. Fresh, K.E. Frick, and T.P. Quinn. 2017. Residency, partial migration, and late egress of subadult Chinook salmon (Oncorhynchus tshawytscha) and coho salmon (O. kisutch) in Puget Sound, Washington. Fishery Bulletin 115(4): 544-555.

Lamperth, J.S., T.P. Quinn, and M.S. Zimmerman. 2017. Levels of stored energy but not marine foraging patterns differentiate seasonal ecotypes of wild and hatchery steelhead (Oncorhynchus mykiss) returning to the Kalama River, Washington. Canadian Journal of Fisheries and Aquatic Sciences 74: 157-167.

Quinn, T.P., M.H. Bond, S.J. Brenkman, R. Paradis, and R.J. Peters. 2017. Re-awakening dormant life history variation: stable isotopes indicate anadromy in bull trout following dam removal on the Elwha River, Washington. Environmental Biology of Fishes. 100: 1659-1671.

Quinn, T.P., C.J. Cunningham, and A.J. Wirsing. 2017. Diverse foraging opportunities drive the functional response of local and landscape-scale bear predation on Pacific salmon. Oecologia. 183: 415-429.

Reed, T.E., E. Eyto, K. O’Higgins, P. Gargan, W. Roche, J. White, N. O’Maoileidigh, T.P. Quinn, and P. McGinnity. 2017. Availability of holding habitat in lakes and rivers affects the incidence of spring (premature) upriver migration by Atlantic salmon. Canadian Journal of Fisheries and Aquatic Sciences. 74: 668-679.

Shaffer, A.J., F. Juanes, T.P. Quinn, D. Parks, T. McBride, J. Michel, C. Naumann, M. Hocking, and C. Byrnes. 2017. Nearshore fish community responses to large scale dam removal: implications for watershed restoration and fish management. Aquatic Sciences.

Sparks, M.M., P.A.H. Westley, J.A. Falke, and T.P. Quinn. Thermal adaptation and phenotypic plasticity in a warming world: Insights from common garden experiments on Alaskan sockeye salmon. Global Change Biology. 23: 5203-5217.

Thornton, E.J., J.J. Duda, and T.P.Quinn. 2017. Influence of species, size and relative abundance on the outcomes of competitive interactions between brook trout and juvenile coho salmon. Ethology Ecology & Evolution 29(2): 157-169.

Tillostson, M.D., and T.P. Quinn. 2017. Climate and conspecific density trigger pre-spawning mortality in sockeye salmon (Oncorhynchus nerka). Fisheries Research 188: 138-148.

Berdahl, A., P.A.H. Westley, S.A. Levin, I.D. Couzin, and T.P. Quinn. 2016. A collective navigation hypothesis for homeward migration in anadromous salmonids. Fish and Fisheries 17: 525-542.

Dennert, A.M., S.L. May-McNally, M.H. Bond, T.P. Quinn, and E.B. Taylor. 2016. Trophic biology and migratory patterns of sympatric Dolly Varden (Salvelinus malma) and Arctic char (Salvelinus alpinus). Canadian Journal of Zoology 94: 529-539.

Hansen, A.G., J.R. Gardner, D.A. Beauchamp, R. Paradis, T.P. Quinn. 2016. Recovery of sockeye salmon in the Elwha River, Washington, after dam removal: dependence of smolt production on the resumption of anadromy by landlocked kokanee. Transactions of the American Fisheries Society. 145: 1303-1317.

Jaecks, T., M.H. Bond, and T.P. Quinn. 2016. Can dietary reliance on Pacific salmon eggs create otolith Sr/Ca signatures that mimic anadromy in resident salmonids? Environmental Biology of Fishes 99: 237-247.

Quinn, T.P., P. McGinnity, and T.E. Reed. 2016. The paradox of “premature migration” by adult anadromous salmonid fishes: patterns and hypotheses. Canadian Journal of Fisheries and Aquatic Sciences 73: 1015-1030.

Tillotson, M.D. and T.P. Quinn. 2016. Beyond correlation in the detection of climate change impacts: testing a mechanistic hypothesis for climatic influence of sockeye salmon (Oncorhynchus nerka) productivity. PLoS ONE 11(4).

Anderson, J.H., P.L. Faulds, K.D. Burton, M.E. Koehler, W.I. Atlas, and T.P. Quinn. 2015. Dispersal and productivity of Chinook (Oncorhynchus tshawytscha) and coho (Oncorhynchus kisutch) salmon colonizing newly accessible habitat. Canadian Journal of Fisheries and Aquatic Sciences 72: 454-465.

Bond, M.H., J.A. Miller, and T.P. Quinn. 2015. Beyond dichotomous life histories in partially migrating populations: cessation of anadromy in a long-lived fish. Ecology.

Goetz, F.A., E. Jeanes, M.E. Moore, and T.P. Quinn. 2015. Comparative migratory behavior and survival of wild and hatchery steelhead (Oncorhynchus mykiss) smolts in riverine, estuarine, and marine habitats of Puget Sound, Washington. Environmental Biology of Fishes 98: 357-375.

Harmon, B.S., Hilborn, R., and T.P. Quinn. 2015. Infection by the cestode parasite Schistocephalus sp. and effects on diet, body condition and survival of sculpins Cottus aleuticus and Cottus cognatus. Journal of Fish Biology 86: 1621-1629.

Hart, L.M., M.H. Bond, S.L. May-McNally, J.A. Miller, and T.P. Quinn. 2015. Use of otolith microchemistry and stable isotopes to investigate the ecology and anadromous migrations of Northern Dolly Varden from the Egegik River, Bristol Bay, Alaska. Environmental Biology of Fishes 98: 1633-1643.

Kendall, N.W., J.R. McMillan, M.R. Sloat, T.W. Buehrens, T.P. Quinn, G.R. Pess, K.V. Kuzishchin, M.M. McClure, and R.W. Zabel. 2015. Anadromy and residency in steelhead and rainbow trout (Oncorhynchus mykiss): a review of the processes and patterns. Canadian Journal of Fisheries and Aquatic Sciences 72: 319-342.

May-McNally, S.L., T.P. Quinn, and E.B. Taylor. Low levels of hybridization between sympatric Arctic char (Salvelinus alpinus) and Dolly Varden char (Salvelinus malma) highlights their genetic distinctiveness and ecological segregation. Ecology and Evolution 5(15): 3031-3045.

May-McNally, S.L., T.P. Quinn, P.J. Woods, and E.B. Taylor. 2015. Evidence for genetic distinction among sympatric ecotypes of Arctic char (Salvelinus alpinus) in south-western Alaskan lakes. Ecology of Freshwater Fish.

McMillan, J.R., G.R. Pess, M. Liermann, S.A. Morley, M.L. McHenry, L.A. Campbell, and T.P. Quinn. 2015. Using redd attributes, fry density, and otolith microchemistry to distinguish the presence of steelhead and rainbow trout in the Elwha River Dam removal project. North American Journal of Fisheries Management 35(5): 1019-1033.

Moore, M.E., B.A. Berejikian, F.A. Goetz, A.G. Berger, S.S. Hodgson, E.J. Connor, T.P. Quinn. 2015. Multi-population analysis of Puget Sound steelhead survival and migration behavior. Marine Ecology Progress Series 537: 217-232.

Quinn, T.P. 2015. Turning class field trips into long-term research: a great idea with few pitfalls. Fisheries 40(2): 65-68.

Quinn, T.P., M.H. Bond, and H.B. Berge. 2015. Use of egg size differences in anadromous (sockeye salmon) and non-anadromous (kokanee) forms of Oncorhynchus nerka to infer ancestral origins of a landlocked population. Ecological Research 30: 547-554.

Smith, J.M., K.L. Fresh, A.N. Kagley, T.P. Quinn. 2015. Ultrasonic telemetry reveals seasonal variation in depth distribution and diel vertical migrations of sub-adult Chinook and coho salmon in Puget Sound. Marine Ecology Progress Series 532: 227-242.

Sprehn, C.G., M.J. Blum, T.P. Quinn, and D.C. Heins. 2015. Landscape genetics of Schistocephalus solidus parasites in threespine stickleback (Gasterosteus aculeatus) from Alaska. PLOS ONE.

Thornton, E.J., R.W. Hardy, and T.P. Quinn. 2015. Experimental determination of the limits of using stable isotopes to distinguish steelhead and rainbow trout offspring. North American Journal of Fisheries Management 35(4): 810-817

Westley, P.A.H., A.H. Dittman, E.J. Ward, and T.P. Quinn. 2015. Signals of climate, conspecific density, and watershed features in patterns of homing and dispersal by Pacific salmon. Ecology.

Adkison, M.D., M.B. Ward, and T.P. Quinn. 2014. Nest site preference and intrasexual competition in female sockeye salmon, Oncorhynchus nerka. Environmental Biology of Fishes 97: 385-399.

Berdahl, A., P.A.H. Westley, S.A. Levin, I.D. Couzin, and T.P. Quinn. 2014. A collective navigation hypothesis for homeward migration in anadromous salmonids. Fish and Fisheries.

Bond, M.H., B.R. Beckman, L. Rohrback, and T.P. Quinn. 2014. Differential growth in estuarine and freshwater habitats indicated by plasma IGF1 concentrations and otolith chemistry in Dolly Varden Salvelinus malma. Journal of Fish Biology 85: 1429-1445.

Bond, M.H., P.A. Crane, W.A. Larson, and T.P. Quinn. 2014. Is isolation by adaptation driving genetic divergence among proximate Dolly Varden char populations? Ecology and Evolution 4(12): 2515-2532.

Buehrens, T.W., P. Kiffney, G.R. Pess, T.R. Bennett, S.M. Naman, G. Brooks, and T.P. Quinn. 2014. Increasing juvenile coho salmon densities during early recolonization have not affected resident coastal cutthroat trout growth, movement, or survival. North American Journal of Fisheries Managment 34: 892-907.

Chamberlin, J.W. and T.P. Quinn. 2014. Effects of natal origin on localized distributions of Chinook salmon, Oncorhynchus tshawytscha, in the marine waters of Puget Sound, Washington. Fisheries Research 153: 113-122.

Jaecks, T. and T.P. Quinn. 2014. Ontogenetic shift to dependence on salmon derived nutrients in Dolly Varden char from Iliamna River, Alaska. Environmental Biology of Fishes 97: 1323-1333.

Kendall, N.W., U. Dieckmann, M. Heino, A.E. Punt, and T.P. Quinn. 2014. Evolution of age and length at maturation of Alaskan salmon under size-selective harvest. Evolutionary Applications 7: 313-322.

McMillan, J.R., G.R. Pess, M.L. McHenry, R. Moses, and T.P. Quinn. 2014. Documentation of unusual, fall spawning by coastal cutthroat trout in the Elwha River system, Washington. Transactions of the American Fisheries Society 143: 1605-1611.

Pess, G.R., T.P. Quinn, S.R. Gephard, and R. Saunders. 2014. Re-colonization of Atlantic and Pacific rivers by anadromous fishes: linkages between life history and the benefits of barrier removal. Reviews in Fish Biology and Fisheries 24: 881-900.

Pess, G.R., T.P. Quinn, D.E. Schindler, and M.C. Liermann. 2014. Freshwater habitat associations between pink (Oncorhynchus gorbuscha), chum (O. keta) and Chinook salmon (O. tshawytscha) in a watershed dominated by sockeye salmon (O. nerka) abundance. Ecology of Freshwater Fish 23: 360-372.

Putman, N.F., M.M. Scanlan, E.J. Billman, J.P. O’Neil, R.B. Couture, T.P. Quinn, K.J. Lohmann, and D.L.G. Noakes. 2014. Inherited magnetic map guides ocean navigation in juvenile Pacific salmon. Current Biology 24: 446-450.

Quinn, T.P., M. Bond, and S. Slater. 2014. Use of stable isotopes and otolith micro-chemistry to evaluate migration in male Chinook salmon, Oncorhynchus tshawytscha, from an Alaskan river. Northwest Science 88(4): 360-366.

Quinn, T.P., C.J. Cunningham, J. Randall, and R. Hilborn. 2014. Can intense predation by bears exert a depensatory effect on recruitment in a Pacific salmon population? Oecologia 176: 445-456.

Quinn, T.P., J.A. Shaffer, J. Brown, N. Harris, C. Byrnes, and P. Crain. 2014. Juvenile Chinook salmon, Oncorhynchus tshawytscha, use of the Elwha River estuary prior to dam removal. Environmental Biology of Fishes 97: 731-740.

Quinn, T.P. A.J. Wirsing, B. Smith, C.J. Cunningham, and J. Ching. 2014. Complementary use of motion-activated cameras and unbaited wire snares for DNA sampling reveals diel and seasonal activity patterns of brown bears (Ursus arctos) foraging on adult sockeye salmon (Oncorhynchus nerka). Canadian Journal of Zoology 92: 893-903.

Rohde, J., K.L. Fresh, and T.P. Quinn. 2014. Factors affecting partial migration in Puget Sound Coho Salmon (Oncorhynchus kisutch). North American Journal of Fisheries Management 34: 559-570.

Taugbøl, A., C. Junge, T.P. Quinn, A. Herland, L.A. Vøllestad. 2014. Genetic and morphometric divergence in threespine stickleback in the Chignik catchment, Alaska. Ecology and Evolution 4: 144-156.

Anderson, J.H., P. Faulds, W.I. Atlas, and T.P. Quinn. 2013. Reproductive success of captively bred and naturally spawned Chinook salmon colonizing newly accessible habitat. Evolutionary Applications 6: 165-179.

Anderson, J.H., G.R. Pess, P.M. Kiffney, T.R. Bennett, P.L. Faulds, W.I. Atlas, and T.P. Quinn. 2013. Dispersal and tributary immigration by juvenile coho salmon contribute to spatial expansion during colonization. Ecology of Freshwater Fish 22: 30-42.

Bond, M.H. and T.P. Quinn. 2013. Patterns and influences on Dolly Varden migratory timing in Chignik Lakes, Alaska, and comparison to populations throughout the Northeastern Pacific and Arctic oceans. Canadian Journal of Fisheries and Aquatic Sciences. 70: 655-665.

Buehrens, T.W., J. Glasgow, C.O. Ostberg, and T.P. Quinn. 2013. Spatial segregation of spawning habitat limits hybridization between sympatric native steelhead and coastal cutthroat trout. Transactions of the American Fisheries Society 142: 221-233.

Cunningham, C.J., M.G. Courage, and T.P. Quinn. 2013. Selecting for the phenotypic optimum: size-related trade-offs between mortality risk and reproductive output in female sockeye salmon. Functional Ecology 27: 1233-1243.

Cunningham, C.J., G.T. Ruggerone, and T.P. Quinn. 2013. Size-selectivity of predation by brown bears depends on the density of their sockeye salmon prey. American Naturalist 181: 663-673.

Essington, T.E., K. Dodd and T.P. Quinn. 2013. Shifts in the estuarine demersal fish community following a fishery closure in Puget Sound, WA. Fishery Bulletin 111: 205-217. 

Goetz, F.A., B. Baker, T. Buehrens, and T.P. Quinn. 2013. Diversity of movements by individual anadromous coastal cutthroat trout Oncorhynchus clarkii clarkii. Journal of Fish Biology 83: 1161-1182.

Kendall, N.W. and T.P. Quinn. 2013. Size-selective fishing affects sex ratios and the opportunity for sexual selection in Alaskan sockeye salmon Oncorhynchus nerka. Oikos 122: 411-420.

Naish, K.A., T.R. Seamons, M.B. Dauer, L. Hauser, and T.P. Quinn. 2013. Relationship between effective population size, inbreeding, and adult fitness-related traits in a steelhead (Oncorhynchus mykiss) population released in the wild. Molecular Ecology 22: 1295-1309.

Putman, N.F., K.J. Lohmann, E.M. Putman, T.P. Quinn, A.P. Klimley, and D.L.G. Noakes. 2013. Evidence for geomagnetic imprinting as a homing mechanism in Pacific salmon. Current Biology 23: 312-316.

Quinn, T.P., N. Harris, J.A. Shaffer, C. Byrnes, and P. Crain. 2013. Juvenile coho salmon, Oncorhynchus kisutch, in the Elwha River estuary prior to dam removal: Seasonal occupancy, size distribution, and comparison to nearby Salt Creek. Transactions of the American Fisheries Society 42: 1058-1066.

Rhode, J., A.N. Kagley, K.L. Fresh, F.A. Goetz, and T.P. Quinn. 2013. Partial migration and diel movement patterns in Puget Sound coho salmon. Transactions of the American Fisheries Society 142: 1615-1628.

Simmons, R.K., T.P. Quinn, L.W. Seeb, D.E. Schindler, and R. Hilborn. 2013. Role of estuarine rearing for sockeye salmon in Alaska (USA). Marine Ecology Progress Series 481: 211–223. 

Simmons, R.K., T.P. Quinn, L.W. Seeb, D.E. Schindler, and R. Hilborn. 2013. Summer emigration and resource acquisition within a shared nursery lake by sockeye salmon (Oncorhynchus nerka) from historically discrete rearing environments. Canadian Journal of Fisheries and Aquatic Sciences 70: 57-63.

Westley, P.A.H., T.P. Quinn and A.H. Dittman. 2013. Rates of straying by hatchery-produced Pacific salmon and steelhead (Oncorhynchus spp.) differ among species, life history types, and populations. Canadian Journal of Fisheries and Aquatic Sciences. 70: 735-746.

Woods, P.J., S. Skúlason, S.S. Snorrason, B.K. Kristjánsson, H.J. Malmquist, and T.P. Quinn. 2013. Intraspecific diversity in Arctic charr, Salvelinus alpinus, in Iceland: I. Detection using mixture models. Evolutionary Ecology Research 14: 973-992.

Woods, P.J., S. Skúlason, S.S. Snorrason, B.K. Kristjánsson, H.J. Malmquist, and T.P. Quinn. 2013. Intraspecific diversity in Arctic charr, Salvelinus alpinus, in Iceland: II. What environmental factors influence resource polymorphism in lakes? Evolutionary Ecology Research 14: 993-1013.

Woods, P.J., D. Young, S. Skúlason, S.S. Snorrason, and T.P. Quinn. 2013. Resource polymorphism and diversity of Arctic charr Salvelinus alpinus in a series of isolated lakes in southwestern Alaska. Journal of Fish Biology 82: 569-587.

Anderson, J.H., P.L. Faulds, W.I. Atlas, and T.P. Quinn. 2012. Reproductive success of captively bred and naturally spawned Chinook salmon colonizing newly accessible habitat. Evolutionary Applications: 1-15.

Andrews, K.S. and T.P. Quinn. 2012. Combining fishing and acoustic monitoring data to evaluate the distribution and movements of spotted ratfish Hydrolagus colliei. Marine Biology159: 769-782.

Gowell, C.P., T.P. Quinn and E.B. Taylor. 2012. Coexistence and origin of trophic ecotypes of pygmy whitefish, Prosopium coulterii, in a southwestern Alaskan lake. Journal of Evolutionary Biology 25: 2432-2448.

Kendall, N.W. and T.P. Quinn. 2012. Quantifying and comparing size-selectivity among Alaskan sockeye salmon fisheries. Ecological Applications 22: 804-816.

Pess, G.R., R. Hilborn, K. Kloehn, and T.P. Quinn. 2012. The influence of population dynamics and environmental conditions on pink salmon re-colonization after barrier removal. Canadian Journal of Fisheries and Aquatic Sciences 69: 970-982.

Quinn, T.P., A.H. Dittman, H. Barrett, C. Cunningham, and M.H. Bond. 2012. Chemosensory responses juvenile coho salmon, Oncorhynchus kisutch, Dolly Varden, Salvelinus malma, and sculpins (Cottus spp.) to eggs and other tissues from adult Pacific salmon. Environmental Biology of Fishes 95: 301-307.

Quinn, T.P., N.W. Kendall, H.B. Rich, Jr., and B.E. Chasco. 2012. Diel vertical movements, and effects of infection by the cestode Schistocephalus solidus on daytime proximity of three-spined sticklebacks (Gasterosteus aculeatus) to the surface of a large Alaskan lake. Oecologia 168: 43-51.

Quinn, T.P., H.B. Rich, Jr., D. Gosse, and N. Schtickzelle. 2012. Population dynamics and asynchrony at fine spatial scales: A case history of sockeye salmon population structure in Alaska. Canadian Journal of Fisheries and Aquatic Sciences 69: 297-306.

Quinn, T.P., T.R. Seamons, and S.P. Johnson. 2012. Stable isotopes of carbon and nitrogen indicate differences in marine ecology between wild and hatchery-produced steelhead trout, Oncorhynchus mykiss. Transactions of the American Fisheries Society 141: 526-532.

Quinn, T.P., C.J. Sergeant, A.H. Beaudreau, and D.A. Beauchamp. 2012. Spatial and temporal patterns of vertical distribution for three planktivorous fishes in Lake Washington. Ecology of Freshwater Fish 21: 337-348.

Seamons, T.R., L. Hauser, K.A. Naish, and T.P. Quinn. 2012. Can interbreeding of wild and artificially propagated animals be prevented by using broodstock selected for a divergent life history? Evolutionary Applications 5: 705-719.

Sharma, R. and T.P. Quinn. 2012. Linkages between life history type and migration pathways in freshwater and marine environments for Chinook salmon, Oncorhynchus tshawytscha. Acta Oecologica 41: 1-13.

Baker, M.R., N.W. Kendall, T.A. Branch, D.E. Schindler, and T.P. Quinn. 2011. Selection due to non-retention mortality in gillnet fisheries for salmon. Evolutionary Applications 4: 429-443.

Carlson, S.M., T.P. Quinn, and A.P. Hendry. 2011. Eco-evolutionary dynamics in Pacific salmon. Heredity 106:438-447.

Chamberlin, J.W., T.E. Essington, J.W. Ferguson, and T.P. Quinn. 2011. The influence of hatchery rearing practices on salmon migratory behavior: Is the tendency of Chinook salmon to remain within Puget Sound affected by size and date of release? Transactions of the American Fisheries Society 140: 1398-1408.

Chamberlin, J.W., A.N. Kagley, K.L. Fresh, and T.P. Quinn. 2011. Movements of yearling Chinook salmon during the first summer in marine waters of Hood Canal, Washington. Transactions of the American Fisheries Society 140:429-439.

Gomez-Uchida, D., M.J. Smith, C. Habicht, J.E Seeb, T.P. Quinn, and L.W. Seeb. Single nucleotide polymorphisms unravel hierarchical divergence and signatures of selection among Alaskan sockeye salmon (Oncorhynchus nerka) populations. BMC Evolutionary Biology.

Kendall, N.W. and T.P. Quinn. 2011. Length and age trends of Chinook salmon in the Nushagak River, Alaska related to commercial and recreational fishery selection and exploitation. Transactions of the American Fisheries Society 140: 611-622.

Kinnison, M.T., T.P. Quinn, and M.J. Unwin. 2011. Correlated contemporary evolution of life history traits in New Zealand Chinook salmon, Oncorhynchus tshawytscha. Heredity 106:448-459.

Krkošek, M., R. Hilborn, R.M. Peterman, and T.P. Quinn. 2011. Cycles, stochasticity, and density dependence in pink salmon population dynamics. Proceeding of the Royal Society of London (B) 278: 2060-2068.

Lin, J.E., R. Hilborn, T.P. Quinn, and L. Hauser. 2011. Self-sustaining populations, population sinks or aggregates of strays: chum (Oncorhynchus keta) and Chinook salmon (O. tshawytscha) in the Wood River system, Alaska. Molecular Ecology 40: 4925-4937.

Pess, G.R., P.M. Kiffney, M.C. Liermann, T.R. Bennett, J.H. Anderson, and T.P. Quinn. 2011. The influences of body size, habitat quality, and competition on the movement and survival of juvenile coho salmon, Oncorhynchus kisutch, during the early stages of stream re-colonization. Transactions of the American Fisheries Society 140: 883-897.

Quinn, T.P., J. Chamberlin, and E.L. Brannon. 2011. Experimental evidence of population-specific spatial distributions of Chinook salmon, Oncorhynchus tshawytscha. Environmental Biology of Fishes 92: 313-322.

Quinn, T.P., T.R. Seamons, L.A. Vøllestad, and E. Duffy. 2011. Effects of growth and reproductive history on the egg size – fecundity tradeoff in steelhead. Transactions of the American Fisheries Society 140: 123-135.

Quinn, T.P., M.J. Unwin and M.T. Kinnison. 2011. Contemporary divergence in migratory timing of naturalized populations of Chinook salmon, Oncorhynchus tshawytscha, in New Zealand. Evolutionary Ecology Research 13: 45-54.

Anderson, J.H., P.L. Faulds, W.I. Atlas, G.R. Pess, and T.P. Quinn. 2010. Selection on breeding date and body size in colonizing coho salmon, Oncorhynchus kisutch. Molec. Ecol. 19:2562-2573.

Armstrong, J.B., D.E. Schindler, K.L. Omori, C.P. Ruff, and T.P. Quinn. 2010. Thermal heterogeneity mediates the effects of pulsed subsidies across a landscape. Ecology 91(5):1445-1454.

Denton, K.P., H.B. Rich, Jr., J.W. Moore, and T.P. Quinn. 2010. The utilization of a Pacific salmon Oncorhynchus nerka subsidy by three populations of charr Salvelinus spp. J. Fish Biol. 77:1006-1023.

Doctor, K.K., R. Hilborn, M. Rowse and T.P. Quinn. 2010. Spatial and temporal patterns of upriver migration by sockeye salmon, Oncorhynchus nerka, populations in the Wood River system, Bristol Bay, Alaska. Transactions of the American Fisheries Society 139: 80-91.

Greene, C.M., J.E. Hall, K.R. Guilbault, and T.P. Quinn. 2010. Improved viability of populations with diverse life history portfolios. Biology Letters 6: 382-386.

Kendall, N.W., H.B. Rich, Jr., L.R. Jensen, and T.P. Quinn. 2010. Climate effects on inter-annual variation in growth of the freshwater mussel, Anodonta beringiana, in Iliamna Lake, Alaska. Freshwat. Biol. 55:2339-2346.

Kendall, N.W., A.H. Dittman, and T.P. Quinn. 2010. Comparative maturation schedules of two Columbia River sockeye salmon, Oncorhynchus nerka, populations. Endangered Species Research 13: 51-61.

Moore, M.E., F.A. Goetz, D.M. Van Doornik, E.P. Tezak, T.P. Quinn, J.J. Reyes-Tomassini, and B.A. Berejikian. 2010. Early marine migration patterns of wild coastal cutthroat trout (Oncorhynchus clarki clarki), steelhead trout (Oncorhynchus mykiss), and their hybrids. PLoS ONE 5(9):e12881.

Reed, T.E., G. Martinek, and T.P. Quinn. 2010. Lake-specific variation in growth, migration timing and survival of juvenile sockeye salmon Oncorhynchus nerka: separating environmental from genetic influences. J. Fish Biol. 77:692-705.

Schindler, D.E., R. Hilborn, B. Chasco, C.P. Boatright, T.P. Quinn, L.A. Rogers, and M.S. Webster. 2010. Population diversity and the portfolio effect in an exploited species. Nature 465:609-613.

Seamons, T.R. and T.P. Quinn. 2010. Sex-specific patterns of lifetime reproductive success in single and repeat breeding steelhead trout (Oncorhynchus mykiss). Behav. Ecol. Sociobiol. 64:505–513.

Westley, P.A.H., D.E. Schindler, G.T. Ruggerone, T.P. Quinn and R. Hilborn. 2010. Natural habitat change, commercial fishing, climate, and dispersal interact to restructure an Alaskan fish metacommunity. Oecologia 163: 471-484.

Woody, C.A., R.M. Hughes, E.J. Wagner, T.P. Quinn, L.H. Roulson, L.M. Martin, and K. Griswold. 2010. The Mining Law of 1872: Change is overdue. Fisheries 35: 321-331.

Carlson, S.M., H.B. Rich, Jr., and T.P. Quinn. 2009. Does variation in selection imposed by bears drive divergence among populations in the size and shape of sockeye salmon? Evolution 63-5:1244-1261.

Dauer, M.B., T.R. Seamons, L. Hauser, T.P. Quinn, and K.A. Naish. 2009. Estimating the ratio of hatchery-produced to wild adult steelhead on the spawning grounds using scale pattern analyses. Trans. Am. Fish. Soc. 138:15-22.

Denton, K.P., H.B. Rich, Jr., and T.P. Quinn. 2009. Diet, movement, and growth of Dolly Varden in response to sockeye salmon subsidies. Trans. Am. Fish. Soc. 138:1207-1219.

Doctor, K.K., and T.P. Quinn. 2009. Potential for adaptation-by-time in sockeye salmon (Oncorhynchus nerka): the interactions of body size and in-stream reproductive life span with date of arrival and breeding location. Can. J. Zool. 87:708-717.

Kendall, N.W., J.J. Hard, and T.P. Quinn. 2009. Quantifying six decades of fishery selection for size and age at maturity in sockeye salmon. Evol. Applic. 2:523-536.

Kendall, N.W. and T.P. Quinn. 2009. Effects of population-specific variation in age and length on fishery selection and exploitation rates of sockeye salmon (Oncorhynchus nerka). Can. J. Fish. Aquat. Sci. 66:896–908.

Quinn, T.P. 2009. Changing themes in the study of diadromous fishes. Pages 823-828 in A. Haro, K.L. Smith, R.A. Rulifson, C.M. Moffitt, R.J. Klauda, M.J. Dadswell, R.A. Cunjak, J.E. Cooper, K.L. Beal, T.S. Avery (eds.), Challenges for Diadromous Fishes in a Dynamic Global Environment. Am. Fish. Soc. Symp. 69:23-42.

Quinn, T.P. 2009. Pacific salmon population structure and dynamics: a perspective from Bristol Bay on life history variation across spatial and temporal scales. Pages 857-871 in CC Krueger, CE Zimmerman (eds), Pacific Salmon: Ecology and Management of Western Alaska’s Populations. Am. Fish. Soc. Symp. 70, Bethesda, Maryland.

Quinn, T.P., S.M. Carlson, S.M. Gende, and H.B. Rich, Jr. 2009. Transportation of Pacific salmon carcasses from streams to riparian forests by bears. Can. J. Zool. 87:195-203.

Quinn, T.P., K. Doctor, N. Kendall, and H.B. Rich, Jr. 2009. Diadromy and the life history of sockeye salmon: nature, nurture, and the hand of man. Pages 23-42 in A. Haro, K.L. Smith, R.A. Rulifson, C.M. Moffitt, R.J. Klauda, M.J. Dadswell, R.A. Cunjak, J.E. Cooper, K.L. Beal, T.S. Avery (eds.), Challenges for Diadromous Fishes in a Dynamic Global Environment. Am. Fish. Soc. Symp. 69:23-42.

Quinn, T.P. and T.R. Seamons. 2009. Tales from scales: old DNA yields insights into contemporary evolutionary processes affecting fishes. Molec. Ecol. 18:2545–2546.

Rich, H.B., Jr., T.P. Quinn, M.D. Scheuerell, and D.E. Schindler. 2009. Climate and intraspecific competition control the growth and life history of juvenile sockeye salmon (Oncorhynchus nerka) in Iliamna Lake, Alaska. Can. J. Fish. Aquat. Sci. 66:238-246.

Seamons, T.R., M.B. Dauer, J. Sneva, and T.P. Quinn. 2009. Use of parentage assignment and DNA genotyping to validate scale analysis for estimating steelhead age and spawning history. N. Am. J. Fish. Mgmt. 29:396-403.

Anderson, J.H., P.M. Kiffney, G.R. Pess, and T.P. Quinn. 2008. Summer distribution and growth of juvenile coho salmon during colonization of newly accessible habitat. Trans. Am. Fish. Soc. 137:772-781.

Crozier, L.G., A.P. Hendry, P.W. Lawson, T.P. Quinn, N.J. Mantua, J. Battin, R.G. Shaw, and R.B. Huey. 2008. Potential responses to climate change in organisms with complex life histories: evolution and plasticity in Pacific salmon. Evol. Applic. 1(2):252-270.

Hauser, D.D.W., C.S. Allen, H.B. Rich, Jr., and T.P. Quinn. 2008. Resident harbor seals (Phoca vitulina) in Iliamna Lake, Alaska: Summer diet and partial consumption of adult sockeye salmon (Oncorhynchus nerka). Aquat. Mamm. 34(3):303-309.

Kinnison, M.T., M.J. Unwin, and T.P. Quinn. 2008. Eco-evolutionary vs. habitat contributions to invasion in salmon: experimental evaluation in the wild. Molec. Ecol. 17:405-414.

Lin, J., T.P. Quinn, R. Hilborn, and L. Hauser. 2008. Fine-scale differentiation between sockeye salmon ecotypes and the effect of phenotype on straying. Heredity 101:341-350.

Lin, J., E. Ziegler, T.P. Quinn, and L. Hauser. 2008. Contrasting patterns of morphological and neutral genetic divergence among geographically proximate populations of sockeye salmon Oncorhynchus nerka in Lake Aleknagik, Alaska. J. Fish Biol. 73:1993-2004.

McLean, J.E., T.R. Seamons, M.B. Dauer, P. Bentzen, and T.P. Quinn. 2008. Variation in reproductive success and effective number of breeders in a hatchery population of steelhead trout (Oncorhynchus mykiss): examination by microsatellite-based parentage analysis. Conserv. Genet. 9:295-304.

Taylor, E.B., E. Lowery, A. Lilliestrale, A. Elz, and T.P. Quinn. 2008. Genetic analysis of sympatric char populations in western Alaska: Arctic char (Salvelinus alpinus) and Dolly Varden (Salvelinus malma) are not two sides of the same coin. J. Evol. Biol. 21:1609-1625.

Westley, P.A.H., S.M. Carlson, and T.P. Quinn. 2008. Among-population variation in adipose fin size parallels the expression of other secondary sexual characteristics in sockeye salmon (Oncorhynchus nerka). Environ. Biol. Fish 81:439-446.

Westley, P.A.H., R. Hilborn, T.P. Quinn, G.T. Ruggerone, and D.E. Schindler. 2008. Long-term changes in rearing habitat and downstream movement by juvenile sockeye salmon (Oncorhynchus nerka) in an interconnected Alaska lake system. Ecol. Freshwat. Fish. 17:443–454.

Allen, C.S., H.B. Rich, Jr, and T.P. Quinn. 2007. Condition-dependent reproductive tactics by large and small anadromous male sockeye salmon Oncorhynchus nerka. J. Fish Biol. 70:1302-1307.

Anderson, J.H. and T.P. Quinn. Movements of adult coho salmon (Oncorhynchus kisutch) during colonization of newly accessible habitat. Can. J. Fish. Aquat. Sci. 64:1143-1154.

Carlson, S.M., R. Hilborn, A.P. Hendry, and T.P. Quinn. 2007. Predation by bears drives senescence in natural populations of salmon. PLoS One 2(12):ed1286.

Carlson, S. and T.P. Quinn. 2007. Ten years of varying lake level and selection on size-at-maturity in sockeye salmon. Ecology 88(10):2620-2629.

Garcia de Leaniz, C., I.A. Fleming, S. Einum, E. Verspoor, W.C. Jordon, S. Consuegra, N. Aubin-Horth, D. Lajus, B.H. Letcher, A.F. Youngson, J.H. Webb, L.A. Vøllestad, B. Villanueva, A. Ferguon, and T.P. Quinn. 2007. A critical review of adaptive genetic variation in Atlantic salmon: implications for conservation. Biol. Rev. 82:173-211.

Naish, K.A., J.E. Taylor, P.S. Levin, T.P. Quinn, J.R. Winton, D. Huppert, and R. Hilborn. 2007. An evaluation of the effects of conservation and fishery enhancement hatcheries on wild populations of salmon. Advanc. Mar. Biol. 53:61-194.

Newell, J.C., K.L. Fresh, and T.P. Quinn. 2007. Arrival patterns and movements of adult sockeye salmon in Lake Washington: implications for management of an urban fishery. N. Am. J. Fish. Mgmt. 27:908-917.

Quinn, T.P., D.M. Eggers, J.H. Clark, and H.B. Rich, Jr. 2007. Density, climate, and the processes of prespawning mortality and egg retention in Pacific salmon (Oncorhynchus spp.). Can. J. Fish. Aquat. Sci. 64:574-582.

Quinn, T.P., S. Hodgson, L. Flynn, R. Hilborn, and D.E. Rogers. 2007. Directional selection by fisheries and the timing of sockeye salmon (Oncorhynchus nerka) migrations. Ecol. Applic. 17(3):731-739.

Schtickzelle, N. and T.P. Quinn. 2007. A metapopulation perspective for salmon and other anadromous fish. Fish and Fisheries 8:297-314.

Seamons, T.R., P. Bentzen, and T.P. Quinn. 2007. DNA parentage analysis reveals inter-annual variation in selection: results from 19 consecutive brood years in steelhead trout. Evol. Ecol. Res. 9(3):409-431.

Hauser, L., T.R. Seamons, M. Dauer, K.A. Naish, and T.P. Quinn. 2006. An empirical verification of population assignment methods with parentage data: hatchery and wild steelhead (Oncorhynchus mykiss) in Forks Creek, Washington, USA. Molec. Ecol. 15:3157-3173.

Hodgson, S., T.P. Quinn, R. Hilborn, R.C. Francis, and D.R. Rogers. 2006. Marine and freshwater climatic factors affecting interannual variation in the timing of return migration to fresh water of sockeye salmon (Oncorhynchus nerka). Fish. Oceanog. 15:1-24.

Johnson, S.P., S.M. Carlson, and T.P. Quinn. 2006. Tooth size and skin thickness in mature sockeye salmon: evidence for habitat constraints and variable investment between the sexes. Ecol. Freshwat. Fish 15:331-338.

Quinn, T.P., P. McGinnity, and T.F. Cross. 2006. Long-term declines in body size and shifts in run timing of Atlantic salmon in Ireland. J. Fish Biol. 68:1713-1730.

Quinn, T.P., I.J. Stewart, and C.P. Boatright. 2006. Experimental evidence of homing to site of incubation by mature sockeye salmon, Oncorhynchus nerka. Anim. Behav. 72:941-949.

Rich, H.B., Jr, S.M. Carlson, B.E. Chasco, K.C. Briggs, and T.P. Quinn. 2006. Movements of male sockeye salmon, Oncorhynchus nerka, on spawning grounds: effects of in-stream residency, density, and body size. Anim. Behav. 71:971-981.

Dickerson, B.R., K.W. Brinck, M.F. Willson, P. Bentzen, and T.P. Quinn. 2005. Relative importance of salmon body size and arrival time at breeding grounds to reproductive success. Ecology 86:347-352.

Dickerson, B.R., M.F. Willson, P. Bentzen, and T.P. Quinn. 2005. Heritability of life history and morphological traits in a wild pink salmon, Oncorhynchus gorbuscha, population assessed by DNA parentage analysis. Trans. Am. Fish. Soc. 134:1323-1328.

Magnuson, J.J. and T.P. Quinn. 2005. Arthur D. Hasler: he showed us the way. (obituary). Environ. Biol. Fish. 74:67-77.

McLean, J.E., P. Bentzen, and T.P. Quinn. 2005. Nonrandom, size- and timing-biased breeding in a hatchery population of steelhead trout. Cons. Biol. 19:446-454.

Meehan, E.P., E.E. Seminet-Reneau, and T.P. Quinn. 2005. Bear predation on Pacific salmon facilitates colonization of carcasses by fly maggots. Am. Midl. Nat. 153:142-151.

Newell, J.C. and T.P. Quinn. 2005. Behavioral thermoregulation by maturing adult sockeye salmon (Oncorhynchus nerka) in a stratified lake prior to spawning. Can. J. Zool. 83:1232-1239.

Quinn, T.P. 2005. Comment: Sperm competition in salmon hatcheries—the need to institutionalize genetically benign spawning protocols. Trans. Am. Fish. Soc. 134:1490-1494.

Quinn, T.P. 2005. The Behavior and Ecology of Pacific Salmon and Trout. University of Washington Press, Seattle. 378 pages.

Quinn, T.P. 2005. Review of Inland Fishes of Washington, by R. Wydoski and R. Whitney. Quarter. Rev. Biol. 80:128-129.

Quinn, T.P., B.R. Dickerson, and L.A. Vøllestad. 2005. Marine survival and distribution patterns of two Puget Sound hatchery populations of coho (Oncorhynchus kisutch) and chinook (O. tshawytscha) salmon. Fish. Res. 76:209-220.

Robins, J.B., C.A. Abrey, T.P. Quinn, and D.E. Rogers. 2005. Lacustrine growth of juvenile pink salmon and a comparison with sympatric sockeye salmon. J. Fish Biol. 66:1671-1680.

Beauchamp, D. A., C. J. Sergeant, M. M. Mazur, J. M. Scheuerell, D. E. Schindler, M. D. Scheuerell, K. L. Fresh, D. E. Seiler, and T. P. Quinn. 2004. Spatial-temporal dynamics of early feeding demand and food supply by sockeye salmon fry in Lake Washington. Trans Am. Fish. Soc. 133:1014-1032.

Boatright, C., T. Quinn and R. Hilborn. 2004. Timing of adult migration and stock structure of sockeye salmon in Bear Lake, Alaska. Trans Am. Fish. Soc. 133: 911-921.

Brannon, E. L., M. S. Powell, T. P. Quinn and A. Talbot. 2004. Population structure of Columbia River basin chinook salmon and steelhead trout. Rev. Fish. Sci. 12:99-232.

Carlson, S.M., H.B. Rich, Jr., and T.P. Quinn. 2004. Reproductive life-span and sources of mortality for alternative male life-history strategies in sockeye salmon, Oncorhynchus nerka. Can. J. Zool. 82:1878–1885.

Dickerson, B. R., M. F. Willson, P. Bentzen and T. P. Quinn. 2004. Size-assortative mating in salmonids: negative evidence for pink salmon in natural conditions. Anim. Behav. 68:381-385.

Gende, S. M. and T. P. Quinn. 2004. The relative importance of prey density and social dominance in determining energy intake by bears feeding on Pacific salmon. Can. J. Zool. 82:75-85.

Gende, S. M., Quinn, T. P., Hilborn, R., Hendry, A. P., and B. Dickerson. 2004. Brown bears selectively kill salmon with higher energy content but only in habitats that facilitate choice. Oikos 104: 518-528.

Gende, S. M., T. P. Quinn, M. F. Willson, R. Heintz, and T. M. Scott. 2004. Magnitude and fate of salmon-derived nutrients and energy in a coastal stream ecosystem. J. Freshwat. Ecol. 19:149-160.

McLean, J. E., Bentzen, P., and Quinn, T. P. 2004. Differential reproductive success of sympatric, naturally spawning hatchery and wild steelhead (Oncorhynchus mykiss). Environm. Biol. Fish. 69:359-369.

Nowak, G. M., R. A. Tabor, E. J. Warner, K. L. Fresh and T. P. Quinn. 2004. Ontogenetic shifts in habitat and diet of cutthroat trout in Lake Washington, Washington. N. Am. J. Fish. Manage. 24:624-635.

Quinn, T. P., L. A. Vøllestad, J. Peterson and V. Gallucci. 2004. Influences of fresh water and marine growth on the egg size – egg number tradeoff in coho and chinook salmon. Trans Am. Fish. Soc. 133:55-65.

Seamons, T. R., P. Bentzen and T. P. Quinn. 2004. The effects of adult length and arrival date on individual reproductive success in wild steelhead trout (Oncorhynchus mykiss). Can. J. Fish. Aqut. Sci. 61:193-204.

Seamons,T. R., P. Bentzen and T. P. Quinn. 2004. The mating system of steelhead (Oncorhynchus mykiss), inferred by molecular analysis of parents and progeny. Environm. Biol. Fish. 69:333-344.

Stewart, I. J., S. M. Carlson, C. P. Boatright, G. B. Buck, and T. P. Quinn. 2004. Site fidelity of spawning sockeye salmon (Oncorhynchus nerka W.) in the presence and absence of olfactory cues. Ecol. Freshwat. Fish. 13:104-110.

Vøllestad, L. A., J. Peterson and T. P. Quinn. 2004. Effects of fresh water and marine growth rates on early maturity in male coho and chinook salmon. Trans Am. Fish. Soc. 133:495-503.

Hilborn, R., T.P. Quinn, D.E. Schindler, D.E. Rogers. 2003. Biocomplexity and fisheries sustainability. Proc. Nat. Acad. Sci. 100:6564-6568.

Kinnison, M.T., M.J. Unwin, T.P. Quinn. 2003. Migratory costs and contemporary evolution of reproductive allocation in male chinook salmon. J. Evolut. Biol. 16:1257-1269.

McLean, J.E., P. Bentzen, T.P. Quinn. 2003. Differential reproductive success of sympatric, naturally-spawning hatchery and wild steelhead trout (Oncorhynchus mykiss) through the adult stage. Can. J. Fish. Aqut. Sci. 60:433-440.

Quinn, T.P. 2003. Migration of Freshwater Fishes. By: Martyn C. Lucas and Etienne Baras. Book Review, in The Quarterly Review of Biology 78:490-491.

Quinn, T.P., S.M. Gende, G.T. Ruggerone, D.E. Rogers. 2003. Density dependent predation by brown bears (Ursus arctos) on sockeye salmon (Oncorhynchus nerka). Can. J. Fish. Aqut. Sci. 60:553-562.

Stewart, I.J., R. Hilborn, T.P. Quinn. 2003. Coherence of observed adult sockeye salmon abundance within and among spawning habitats in the Kvichak River watershed. Alaska Fish. Res. Bull. 10:28-41.

Stewart, I.S., T.P. Quinn, P. Bentzen. 2003. Evidence for fine-scale natal homing among island beach spawning sockeye salmon, Oncorhynchus nerka. Environm. Biol. Fish. 67:77-85.

Unwin, M.J., M.T. Kinnison, N.C. Boustead, T.P. Quinn. 2003. Genetic control over survival in Pacific salmon (Oncorhynchus spp.): experimental evidence between and within populations of New Zealand chinook salmon (O. tshawytscha). Can. J. Fish. Aqut. Sci. 60:1-11.

Vøllestad, L.A., T.P. Quinn. 2003. Trade-off between growth rate and aggression in juvenile coho salmon Oncorhynchus kisutch. Anim. Behav. 66:561-568.

Dickerson, B.R., M.F. Willson, T.P. Quinn. 2002. Body size, arrival date, and reproductive success of pink salmon, Oncorhynchus gorbuscha. Ethol., Ecol. Evol. 14:29-44.

Hodgson, S., T.p. Quinn. 2002. The timing of adult sockeye salmon migration into fresh water: adaptations by populations to prevailing thermal regimes. Can. J. Zool. 80:542-555.

Kinnison, M.T., P. Bentzen, M.J. Unwin, T.P. Quinn. 2002. Reconstructing recent divergence: evaluating non-equilibrium population structure in New Zealand chinook salmon. Molec. Ecol. 11:739-754.

Nowak, G.M., T.P. Quinn. 2002. Diel and seasonal patterns of horizontal and vertical movements of telemetered cutthroat trout in Lake Washington, Washington. Trans. Am. Fish. Soc. 131:452-462.

Quinn, T.P., J.A. Peterson, V. Gallucci, W.K. Hershberger, E.L. Brannon. 2002. Artificial selection and environmental change: countervailing factors affecting the timing of spawning by coho and chinook salmon. Trans. Am. Fish. Soc. 131:591-598.

Robards, M.D., T.P. Quinn. 2002. The migratory timing of adult summer-run steelhead in the Columbia River over six decades of environmental change. Trans. Am. Fish. Soc. 131:523-536.

Barry, T.P., M.J. Unwin, J.A. Malison, T.P. Quinn. 2001. Free and total cortisol levels in semelparous and iteroparous chinook salmon. J. Fish Biol. 59:1673-1676.

Bentzen, P., J.B. Olsen, J. McLean, T.R. Seamons, and T.P. Quinn. Kinship analysis of Pacific salmon: insights into mating, homing and timing of reproduction. Journal of Heredity 92: 127-136.

Courtenay, S.C., T.P. Quinn, H.M.C. Dupuis, C. Groot and P.A. Larkin. Discrimination of family-specific odours by juvenile coho salmon (Oncorhynchus kisutch): roles of learning and odour concentration. Journal of Fish Biology 58: 107-125.

Gende, S.M., T.P. Quinn and M.F. Willson. Consumption choice by bears feeding on salmon. Oecologia 127: 372-382.

Hendry, A.P., O.K. Berg and T.P. Quinn. Choice of location and time of breeding in salmon: causes and consequences. Oikos 93: 407-418.

Kahler, T.H., P. Roni, T.P. Quinn. 2001. Summer movement and growth of juvenile anadromous salmonids in small western Washington streams. Can. J. Fish. Aquat. Sci. 58:1947-1956.

Kinnison, M.T., M.J. Unwin, A.P. Hendry, and T.P. Quinn. Migratory costs and the evolution of egg size and number allocation in new and indigenous salmon populations. Evolution 55: 1656-1667.

Mackey, G., J.E. McLean and T. P. Quinn. 2001. Comparisons of run timing, spatial distribution, and length of wild and newly-established hatchery populations of steelhead in Forks Creek, Washington. North American Journal of Fisheries Management 21:917-937.

Quinn, T.P. 2001. Review of: Sustainable Fisheries Management: Pacific salmon. Edited by E.E. Knutson, C.R. Steward, D.D. MacDonald, J.E. Williams, D.W. Reiser. Copeia 2001:1174-1175.

Quinn, T.P. and G.B. Buck. 2001. Size- and sex-selective mortality of adult sockeye salmon: bears, gulls, and fish out of water. Trans. Am. Fish. Soc. 130:995-1005.

Quinn, T.P., A.E. Hendry, G.B. Buck. 2001. Balancing natural and sexual selection in sockeye salmon: interactions between body size, reproductive opportunity and vulnerability to predation by bears. Evol. Ecol. Res. 3:917–937.

Quinn, T.P., M.T. Kinnison, M.J. Unwin. 2001. Evolution of chinook salmon (Oncorhynchus tshawytscha) populations in New Zealand: pattern, rate, and process. Genetica 112-113:493-513.

Quinn, T.P., L. Wetzel, S. Bishop, K. Overberg, D.E. Rogers. 2001. Influence of breeding habitat on bear predation and age at maturity and sexual dimorphism of sockeye salmon populations. Can. J. Zool. 79:1782-1793. 

Roni, P. and T.P. Quinn. 2001. Density and size of juvenile salmonids in response to placement of large woody debris in western Oregon and Washington streams. Canadian Journal of Fisheries and Aquatic Sciences 58: 282-292.

Roni, P. and T. P. Quinn. 2001. Effects of wood placement on movements of trout and juvenile coho salmon in natural and artificial stream channels. Transactions of the American Fisheries Society 130: 675-685. 

Essington, T.E., T.P. Quinn and V.E. Ewert. Intra- and interspecific competition and the reproductive success of sympatric Pacific salmon. Canadian Journal of Fisheries and Aquatic Sciences 57: 205-213.

Hendry, A.P., J.K. Wenburg, P. Bentzen, E. C. Volk and T. P. Quinn Rapid evolution of reproductive isolation in the wild: evidence from introduced salmon. Science 290: 516-518.

Quinn, T.P. and G.B. Buck. Scavenging by brown bears, Ursus arctos, and glaucous-winged gulls, Larus glaucescens, on adult sockeye salmon, Oncorhynchus nerka. Canadian Field-Naturalist 114: 217-223.

Quinn, T.P., M.J. Unwin, and M.T. Kinnison. Evolution of temporal isolation in the wild: genetic divergence in timing of migration and breeding by introduced chinook salmon populations. Evolution 54: 1372-1385.

Scholz, N.L., N.K. Truelove, B.L. French, B.A. Berejekian, T.P. Quinn, E. Casillas and T.K. Collier. Diazinon disrupts antipredator and homing behavior in chinook salmon (Oncorhynchus tshawytscha). Canadian Journal of Fisheries and Aquatic Sciences 57: 1911-1918.

Schuett-Hames, D.E., N.P. Peterson, R. Conrad and T.P. Quinn. Patterns of gravel scour and fill after spawning by chum salmon in a western Washington stream. North American Journal of Fisheries Management 20: 610-617.

Unwin, M.J., T.P. Quinn, M.T. Kinnison and N.C. Boustead. Divergence in juvenile growth and life history in two recently colonized and partially isolated chinook salmon populations. Journal of Fish Biology 57: 943-960

Candy, J.R. and T.P. Quinn. Behavior of adult chinook salmon (Oncorhynchus tshawytscha) in British Columbia coastal waters determined from ultrasonic telemetry. Canadian Journal of Zoology 77: 1161-1169.

Griffith, J.N., A.P. Hendry and T.P. Quinn. Exploratory behavior or straying: adult sockeye salmon (Oncorhynchus nerka) entering a non-natal hatchery. Fishery Bulletin 97: 713-716.

Hendry, A.P., O.K. Berg and T.P. Quinn. Breeding date, life history, and energy allocation in a population of sockeye salmon (Oncorhynchus nerka). Oikos 85: 499-514.

Montgomery, D.R., E.M. Beamer, G. Pess and T.P. Quinn. Channel type and salmon spawning distribution. Canadian Journal of Fisheries and Aquatic Sciences 56:377-387.

Quinn, T.P. Variation in Pacific salmon reproductive behaviour associated with species, sex and levels of competition. Behaviour 136: 179-204.

Quinn, T.P. Revisiting the stock concept in Pacific salmon: insights from Alaska and New Zealand. Northwest Science 73: 312-324.

Quinn, T.P. and M.T. Kinnison. Size-selective and sex-selective predation by brown bears on sockeye salmon. Oecologia 121: 273-282.

Quinn, T.P., E.C. Volk and A.P. Hendry. Natural otolith microstructure patterns reveal precise homing to natal incubation sites by sockeye salmon (Oncorhynchus nerka). Canadian Journal of Zoology 77: 766-775.

Rhodes, J.S. and T.P. Quinn. Comparative performance of genetically similar hatchery and naturally-reared juvenile coho salmon in streams. North American Journal of Fisheries Management 19: 670-677.

Steen, R.P. and T.P. Quinn. Egg burial depth by sockeye salmon (Oncorhynchus nerka): implications for survival of embryos and natural selection on female body size. Canadian Journal of Zoology 77: 836-841.

Unwin, M.J., M.T. Kinnison and T. P. Quinn. Exceptions to semelparity: postmaturation survival, morphology and energetics of male chinook salmon (Oncorhynchus tshawytscha). Canadian Journal of Fisheries and Aquatic Sciences 56: 1172-1181.

Dittman, A.H., T.P. Quinn and E.C. Volk. Is the distribution, growth and survival of juvenile salmonids sex-biased? Negative results for coho salmon in an experimental stream channel. Journal of Fish Biology 53: 1360-1364.

Hansen, L.P. and T.P. Quinn. The marine phase of the Atlantic salmon life cycle, with comparisons to Pacific salmon. Canadian Journal of Fisheries and Aquatic Sciences (Supplement 1): 104-118.

Kinnison, M., M. Unwin, N. Boustead and T. Quinn. Population-specific variation in body dimensions of adult chinook salmon (Oncorhynchus tshawytscha) from New Zealand and a related Sacramento River population, 90 years after introduction. Canadian Journal of Fisheries and Aquatic Sciences 55: 554-563.

Kinnison, M.T., M.J. Unwin, W.K. Hershberger and T.P. Quinn. Egg size, fecundity and development rate of two New Zealand chinook salmon (Oncorhynchus tshawytscha) populations, with a comparison to their ancestral Sacramento River population. Canadian Journal of Fisheries and Aquatic Sciences 55: 1946-1953.

Marschall, E.A., D.A. Roff, T.P. Quinn, J.A. Hutchings, N.B. Metcalfe, T.A. Bakke, R.L. Saunders and L. Poff. A framework for understanding Atlantic salmon life history. Canadian Journal of Fisheries and Aquatic Sciences (Supplement) 55(Supplement 1):48-58.

McCormick, S.D., Hansen, L. P., Quinn, T. P., and Saunders, R. L. Movement, migration and smolting of Atlantic salmon. Canadian Journal of Fisheries and Aquatic Sciences (Supplement 1) 55: 77-92.

McPhee, M.V. and T.P. Quinn. Factors affecting the duration of nest defense and reproductive lifespan of female sockeye salmon, Oncorhynchus nerka. Environmental Biology of Fishes 51: 369-375.

Quinn, T.P. and M.V. McPhee. Effects of senescence and density on the aggression of adult female sockeye salmon. Journal of Fish Biology 52: 1295-12.

Quinn, T.P., E. Graynoth, C.C. Wood and C.J. Foote. Genotypic and phenotypic divergence of sockeye salmon in New Zealand from their ancestral British Columbia populations. Transactions of the American Fisheries Society 127: 517-534.

Rhodes, J.S. and T.P. Quinn. Factors affecting the outcome of territorial contests between hatchery and naturally reared coho salmon parr in the laboratory. Journal of Fish Biology 53: 1220-1230.

Spidle, A.P., T.P. Quinn and P. Bentzen. Sex-biased marine survival and growth in a population of coho salmon (Oncorhynchus kisutch). Journal of Fish Biology 52: 907-915.

Berg, O.K., M.D. Adkison and T.P. Quinn. Bilateral asymmetry, sexual dimorphism, and nematode parasites in mature male sockeye salmon Oncorhynchus nerka. Northwest Science 71: 305-312.

Courtenay, S.C., T.P. Quinn, H.M.C. Dupuis, C. Groot and P.A. Larkin Factors affecting the recognition of population-specific odours by juvenile coho salmon (Oncorhynchus kisutch). Journal of Fish Biology 50: 1042-1060.

Dittman, A. H., T. P. Quinn, G. A. Nevitt, B. Hacker and D. R. Storm. Sensitization of olfactory guanylyl cyclase to a specific imprinted odorant in coho salmon. Neuron 19: 381-389.

Hendry, A.P. and T.P. Quinn. Variation in adult life history and morphology among Lake Washington sockeye salmon (Oncorhynchus nerka) populations, in relation to habitat features and ancestral affinities. Canadian Journal of Fisheries and Aquatic Sciences 54: 75-84.

Quinn, T.P. Review of “Migration: The Biology of Life on the Move” by Hugh Dingle. Quarterly Review of Biology 72: 232.

Quinn, T.P., S. Hodgson and C. Peven. Temperature, flow and the migration of adult sockeye salmon (Oncorhynchus nerka) in the Columbia River. Canadian Journal of Fisheries and Aquatic Sciences 54: 1349-1360.

Berejekian, B.A., S.B. Mathews and T.P. Quinn. The effects of hatchery and wild ancestry and rearing environments on the development of agonistic behavior in steelhead trout (Oncorhynchus mykiss) fry. Canadian Journal of Fisheries and Aquatic Sciences 53: 2004-2014.

Candy, J.R., E.W. Carter, T.P. Quinn and B.R. Riddell. Adult chinook salmon behavior and survival after catch and release from purse-seine vessels in Johnstone Strait, British Columbia. North American Journal of Fisheries Management 16: 521-529.

Dittman, A.H. and T.P. Quinn. Homing in Pacific salmon: mechanisms and ecological basis. Journal of Experimental Biology 199: 83-91.

Dittman, A.H., T.P. Quinn and G.A. Nevitt. Timing of imprinting to natural and artificial odors by coho salmon (Oncorhynchus kisutch). Canadian Journal of Fisheries and Aquatic Sciences 53: 434-442.

Hawkins, D.K. and T.P. Quinn. Critical swimming velocity and associated morphology of juvenile coastal cutthroat trout (Oncorhynchus clarki clarki), steelhead trout (O. mykiss) and their hybrids. Canadian Journal of Fisheries and Aquatic Sciences 53: 1487-1496.

Hendry, A.P., T.P. Quinn and F.M. Utter. Genetic evidence for the persistence and divergence of native and introduced populations of sockeye salmon (Oncorhynchus nerka) within Lake Washington, WA. Canadian Journal of Fisheries and Aquatic Sciences 53: 823-832.

Montgomery, D.R., J.M. Buffington, N.P. Peterson, D. Schuett-Hames and T.P. Quinn. Stream-bed scour, egg burial depths, and the influence of salmonid spawning on bed surface mobility and embryo survival. Canadian Journal of Fisheries and Aquatic Sciences 53: 1061-1070.

Peterson, N.P. and T.P. Quinn. Persistence of egg pocket architecture in redds of chum salmon (Oncorhynchus keta). Environmental Biology of Fishes 46: 243-253.

Peterson, N.P. and T.P. Quinn. Spatial and temporal variation in dissolved oxygen in natural egg pockets of chum salmon, Oncorhynchus keta (Walbaum), in Kennedy Creek, Washington. Journal of Fish Biology 48: 131-143.

Quinn, T.P. and D.J. Adams. Environmental changes affecting the migratory timing of American shad and sockeye salmon. Ecology 77: 1151-1162.

Quinn, T.P., M.D. Adkison and M.B. Ward. Behavioral tactics of male sockeye salmon (Oncorhynchus nerka) under varying operational sex ratios. Ethology 102: 304-322.

Quinn, T.P., J.L. Nielsen, C. Gan, M.J. Unwin, R. Wilmot, C. Guthrie, and F.M. Utter. Origin and genetic structure of chinook salmon (Oncorhynchus tshawytscha) transplanted from California to New Zealand: Allozyme and mtDNA evidence. Fishery Bulletin 94: 506-521.

Quinn, T.P., N.P. Peterson and S. Spalding. Effects of instream brush on juvenile coho salmon: Response to comment by C.C. Coutant. Transactions of the American Fisheries Society 125: 151-153.

Quinn, T.P. and N.P. Peterson. The influence of habitat complexity and fish size on over-winter survival and growth of individually-marked juvenile coho salmon (Oncorhynchus kisutch) in Big Beef Creek, Washington. Canadian Journal of Fisheries and Aquatic Sciences 53: 1555-1564.

Adkison, M.D., T.P. Quinn and O. Rutten. An inexpensive, non-disruptive method of in situ dart tagging for visual recognition of fish underwater. North American Journal of Fisheries Management 15: 507-511.

Berg, O.K., C.J. Foote and T.P. Quinn. Fish age, nematode, Philonema oncorhynchi, infection and development of sexual dimorphism by the adult male sockeye salmon, Oncorhynchus nerka, in western Alaska. Canadian Journal of Zoology 73: 1999-2004.

Hendry, A.P., F.E. Leonetti and T.P. Quinn. Spatial and temporal isolating mechanisms: the formation of discrete breeding aggregations of sockeye salmon (Oncorhynchus nerka). Canadian Journal of Zoology 73: 339-352.

Konecki, J.T., C.A. Woody and T.P. Quinn. Critical thermal maxima of coho salmon (Oncorhynchus kisutch) fry under field and laboratory acclimation regimes. Canadian Journal of Zoology 73: 993-996.

Konecki, J.T., C.A. Woody and T.P. Quinn. Influence of temperature on incubation rates of coho salmon (Oncorhynchus kisutch) from ten Washington populations. Northwest Science 69: 126-132.

Konecki, J.T., C.A. Woody and T.P. Quinn. Temperature preference in two populations of juvenile coho salmon (Oncorhynchus kisutch). Environmental Biology of Fishes 44: 417-421.

Lonzarich, D.G. and T.P. Quinn. Experimental evidence for the effect of depth and structure on the distribution, growth and survival of stream fishes. Canadian Journal of Zoology 73: 2223-2230.

Pascual, M.A., T.P. Quinn and H. Fuss. Factors affecting the homing of Columbia River hatchery-produced fall chinook salmon. Transactions of the American Fisheries Society 124: 308-320.

Quinn, T.P. Review of Cone, J. A Common Fate: Endangered Salmon and the People of the Pacific Northwest. Illahee, Journal of the Northwest Environment 11: 106-107.

Quinn, T.P., A.P. Hendry and L.A. Wetzel. The influence of life history trade-offs and the size of incubation gravels on egg size variation in sockeye salmon (Oncorhynchus nerka). Oikos 74: 425-438.

Roni, P. and T.P. Quinn. Geographic variation in size and age of North American chinook salmon (Oncorhynchus tshawytscha). North American Journal of Fisheries Management 15: 325-345.

Spalding, S., N.P. Peterson and T.P. Quinn. Summer distribution, survival and growth of juvenile coho salmon Oncorhynchus kisutch under varying experimental conditions of brushy in-stream cover. Transactions of the American Fisheries Society 124: 124-130.

Warner, E.J. and T.P. Quinn. The horizontal and vertical movements of telemetered rainbow trout (Oncorhynchus mykiss) in Lake Washington. Canadian Journal of Zoology 73: 146-153.

Dittman, A.H. and T.P. Quinn. Avoidance of a putative pheromone, 17a,20b-dihydroxy-4-pregnene-3-one, by precociously mature chinook salmon (Oncorhynchus tshawytscha). Canadian Journal of Zoology 72: 215-219.

Dittman, A.H., T.P. Quinn, W.W. Dickhoff and D.A. Larsen. Interactions between novel water, thyroxine and olfactory imprinting in underyearling coho salmon (Oncorhynchus kisutch, Walbaum). Aquaculture and Fisheries Management 25 (Supplement 2): 157-169.

Nevitt, G.A., A.H. Dittman, T.P. Quinn and W.J. Moody, Jr. Evidence for a peripheral olfactory memory in imprinted salmon. Proceedings of the National Academy of Science. 91: 4288-4292.

Pascual, M.A. and T.P. Quinn. Geographical patterns of straying of fall chinook salmon (Oncorhynchus tshawytscha) from Columbia River (U.S.A.) hatcheries. Aquaculture and Fisheries Management 25 (Supplement 2): 17-30.

Peterson, N.P., E.F. Prentice and T.P. Quinn. Comparison of sequential coded-wire and passive integrated transponder tags for assessing overwinter growth and survival of juvenile coho salmon. North American Journal of Fisheries Management 14: 870-873.

Quinn, T.P. Anthropogenic influences on fish populations of the Georgia Basin. Part I: Salmonids. In: Review of the Marine Environment and Biota of Strait of Georgia, Puget Sound and Juan de Fuca Strait. Canadian Technical Report of Fisheries and Aquatic Sciences 1948: 219-229.

Quinn, T.P. How do sharks orient at sea? Trends in Ecology and Evolution 9: 277-278.

Quinn, T.P. Review of Cloud, J.G. and Thorgaard, G.H., Genetic Conservation of Salmonid Fishes. Animal Behavior 48: 1489-1490.

Quinn, T.P., A.H. Dittman, N.P. Peterson and E.C. Volk. Distribution, survival and growth of sibling groups of juvenile coho salmon, Oncorhynchus kisutch, in an experimental stream channel. Canadian Journal of Zoology 72: 2119-2123.

Quinn, T. P. and C. J. Foote. The effects of body size and sexual dimorphism on the reproductive behaviour of sockeye salmon (Oncorhynchus nerka). Animal Behaviour 48: 751-761.

Varnavskaya, N.V., C.C. Wood, R.J. Everett, R.L. Wilmot, V.S. Varnavsky. V.V. Midanaya and T.P. Quinn. Genetic differentiation of subpopulations of sockeye salmon (Oncorhynchus nerka) within lakes of Alaska, British Columbia and Kamchatka, Russia. Canadian Journal of Fisheries and Aquatic Sciences 51 (Supplement 1): 147-157.

Blair, G.R., D.E. Rogers and T.P. Quinn. Variation in life history characteristics and morphology of sockeye salmon (Oncorhynchus nerka) in the Kvichak River system, Bristol Bay, Alaska. Transactions of the American Fisheries Society 122: 550-559.

Olson, A.F. and T.P. Quinn. Vertical and horizontal movements of adult chinook salmon, Oncorhynchus tshawytscha, in the Columbia River estuary. Fishery Bulletin 91: 171-178.

Quinn, T.P. A review of homing and straying of wild and hatchery-produced salmon. Fisheries Research 18: 29-44.

Quinn, T.P. Book Review of Pearcy, W.G. Ocean Ecology of North Pacific Salmonids. Transactions of the American Fisheries Society 122: 513-514.

Quinn, T. P. and M. J. Unwin. Life history patterns of New Zealand chinook salmon (Oncorhynchus tshawytscha) populations. Canadian Journal of Fisheries and Aquatic Sciences 50: 1414-1421.

Unwin, M. J. and T. P. Quinn. Homing and straying patterns of chinook salmon (Oncorhynchus tshawytscha) from a New Zealand hatchery: spatial distribution of strays and effects of release date. Canadian Journal of Fisheries and Aquatic Sciences 50: 1168-1175.

Bisson, P.A., T.P. Quinn, G.H. Reeves and S.V. Gregory. Best management practices, cumulative effects, and long-term trends in fish abundance in Pacific Northwest river systems. In: Watershed Management. R.J. Naiman (editor). Springer-Verlag, pp.189-232.

Quinn, T.P. Book Review of Groot, C. and L. Margolis (editors) Pacific Salmon Life Histories. Copeia 1992: 927-928.

Quinn, T. P. and G.R. Blair. Morphological changes in senescing adult male sockeye salmon (Oncorhynchus nerka Walbaum). Journal of Fish Biology 41: 1045-1047.

Quinn, T P. and S. Bloomberg. Fecundity of chinook salmon (Oncorhynchus tshawytscha) from the Waitaki and Rakaia rivers. New Zealand Journal of Marine and Freshwater Research 26: 429-434.

Quinn, T.P. and A.H. Dittman. Fishes. In: Animal Homing. F. Papi (editor). Chapman and Hall, pp.145-211.

Berman, C.H. and T.P. Quinn. Behavioural thermoregulation and homing by spring chinook salmon, Oncorhynchus tshawytscha (Walbaum), in the Yakima River. Journal of Fish Biology 39: 301-312.

Blair, G.R. and T.P. Quinn. Homing and spawning site selection by sockeye salmon (Oncorhynchus nerka) in Iliamna Lake, Alaska. Canadian Journal of Zoology 69: 176-181.

Moser, M.L., A.F. Olson and T.P. Quinn. Riverine and estuarine migratory behavior of coho salmon (Oncorhynchus kisutch) smolts. Canadian Journal of Fisheries and Aquatic Sciences 48: 1670-1678.

Pascual, M. A. and T. P. Quinn. Evaluation of alternative models of the coastal migration of adult Fraser River sockeye salmon (Oncorhynchus nerka). Canadian Journal of Fisheries and Aquatic Sciences 48: 799-810.

Quinn, T.P. Models of Pacific salmon orientation and navigation on the open ocean. Journal of Theoretical Biology 150: 539-545.

Quinn, T.P. and R.D. Brodeur. Intra-specific variations in the movement patterns of marine animals. American Zoologist 31: 231-241.

Quinn, T.P., R.S. Nemeth and D. O. McIsaac. Patterns of homing and straying by fall chinook salmon in the lower Columbia River. Transactions of the American Fisheries Society 120: 150-156.

Brannon, E.L. and T.P. Quinn. A field test of the pheromone hypothesis for homing by Pacific salmon. Journal of Chemical Ecology 16: 603-609.

Matthews, K.R., T.P. Quinn and B.S. Miller. The application of ultrasonic transmitters to demersal rockfish movement research on shallow rocky reefs in Puget Sound, Washington. American Fisheries Society Symposium 7: 375-379.

Moser, M.L., A.F. Olson and T.P. Quinn. Effects of dummy ultrasonic transmitters on juvenile coho salmon. American Fisheries Society Symposium 7: 353-356.

Quinn, T.P. Current controversies in the study of salmon homing. Ethology, Ecology and Evolution 2: 49-63.

Quinn, T. P. Migratory behavior of Pacific salmon in estuaries: recent results with ultrasonic telemetry. In: C. A. Simenstad (editor), Effects of Dredging on Anadromous Pacific Coast Fishes. Washington Sea Grant, Seattle, Washington, pp. 13-25.

Quinn, T.P. and A.H. Dittman. Pacific salmon migrations and homing: mechanisms and adaptive significance. Trends in Ecology and Evolution 5: 174-177.

Ruggerone, G.T., T.P. Quinn, I.A. McGregor and T.S. Wilkinson. Horizontal and vertical movements of adult steelhead trout, Oncorhynchus mykiss, in Dean and Fisher channels, British Columbia. Canadian Journal of Fisheries and Aquatic Sciences 47: 1963-1969.

Brannon, E.L. and T.P. Quinn. Odor cues used by homing coho salmon. In: E.L. Brannon and B. Jonsson (editors), Salmon and Trout Migratory Behavior Symposium. pp. 30-34.

Moser, M.L., A.F. Olson and T.P. Quinn. Migratory behavior of coho salmon smolts (Oncorhynchus kisutch) in Grays Harbor, Washington. Proceedings of the 1988 Chinook and Coho Salmon Workshop. pp. 220-226.

Quinn, T.P. Current controversies in the study of marine migration. Review of Herrnkind, W.F. and A.B. Thistle (eds.) Signposts in the Sea. Environmental Biology of Fishes 26: 311-313. Quinn, T.P. Review of McDowall, R.M., Diadromy in Fishes. 1989. Quarterly Review of Biology 64: 358-359.

Quinn, T.P. and S. Courtenay. Intraspecific chemosensory discrimination in salmonid fishes: Alternative explanations. In: E.L. Brannon and B. Jonsson (editors), Salmon and Trout Migratory Behavior Symposium. pp. 35-41. Quinn, T.P., E.L. Brannon and A.H. Dittman. Spatial aspects of imprinting and homing by coho salmon (Oncorhynchus kisutch). Fishery Bulletin 87: 769-774.

Quinn, T.P. and J.T. Light. Occurrence of threespine sticklebacks (Gasterosteus aculeatus) in the open North Pacific Ocean: migration or drift? Canadian Journal of Zoology 67: 2850-2852.

Quinn, T.P., B.A. terHart and C. Groot. Migratory orientation and vertical movements of homing adult sockeye salmon (Oncorhynchus nerka) in coastal waters. Animal Behaviour 37: 587-599.

McIsaac, D.O. and T.P. Quinn. Evidence for a hereditary component in homing behavior of chinook salmon (Oncorhynchus tshawytscha). Canadian Journal of Fisheries and Aquatic Sciences 45: 2201-2205.

Quinn, T.P. Estimated swimming speeds of migrating adult sockeye salmon. Canadian Journal of Zoology 66: 2160-2163.

Quinn, T.P., A.F. Olson and J.T. Konecki. Effects of anaesthesia on the chemosensory behaviour of Pacific salmon. Journal of Fish Biology 33: 637-641.

Walker, M.M., T.P. Quinn, J.L. Kirschvink and C. Groot. Production of single-domain magnetite throughout life by sockeye salmon, Oncorhynchus nerka. Journal of Experimental Biology 140: 51-63.

Matthews, K.R., B.S. Miller and T.P. Quinn. Movement studies of nearshore demersal rockfishes in Puget Sound, Washington. Proceedings of the Lowell Wakefield International Rockfish Symposium, Anchorage, Alaska, pp. 63-72.

Groot, C. and T.P. Quinn. The homing migration of sockeye salmon to the Fraser River. Fishery Bulletin 85: 455-469.

Quinn, T.P. and W.C. Leggett. Perspectives on the marine migrations of diadromous fishes. American Fisheries Society Symposium 1: 377-388.

Quinn, T.P. and R.F. Tallman. Seasonal environmental predictability and homing in riverine fishes. Environmental Biology of Fishes 18: 155-159.

Quinn, T.P. and B.A. terHart. Movements of adult sockeye salmon (Oncorhynchus nerka) in British Columbia coastal waters in relation to temperature and salinity stratification: Ultrasonic telemetry results. In: H.D. Smith, L. Margolis and C. C. Wood (editors), Sockeye Salmon (Oncorhynchus nerka) Population Biology and Future Management. Canadian Special Publication in Fisheries and Aquatic Sciences 96: 61-77.

Quinn, T.P., C.C. Wood, L. Margolis, B. Riddell and K.D. Hyatt. Homing in wild sockeye salmon (Oncorhynchus nerka) populations as inferred from differences in parasite prevalence and allozyme allele frequencies. Canadian Journal of Fisheries and Aquatic Sciences 44: 1963-1971.

Brannon, E.L., T.P. Quinn, R.P. Whitman, A.E. Nevissi, R.E. Nakatani and C. D. McAuliffe. The homing of adult chinook salmon after brief exposure to whole and dispersed crude oil. Transactions of the American Fisheries Society 115: 823-827.

Groot, C., T.P. Quinn and T.J. Hara. Responses of migrating adult sockeye salmon (Oncorhynchus nerka) to population-specific odors. Canadian Journal of Zoology 64: 926-932.

Quinn, T.P. Review of McKeown, B.A., Fish Migration, and Smith, R.J.F., The Control of Fish Migration. Copeia 1986: 268-269.

Quinn, T.P., K.D. Cooke and G. Ellis. The responses of adult sockeye salmon (Oncorhynchus nerka) to a commercial purse seine. Canadian Technical Report of Fisheries and Aquatic Sciences 1511: 13 p.

Quinn, T.P. and T.J. Hara. Sibling recognition and olfactory sensitivity in juvenile coho salmon (Oncorhynchus kisutch). Canadian Journal of Zoology 64: 921-925.

Quinn, T.P. and G.M Tolson. Evidence of chemically mediated population recognition in coho salmon (Oncorhynchus kisutch). Canadian Journal of Zoology 64: 84-87.

Quinn, T.P. Homing and the evolution of sockeye salmon, Oncorhynchus nerka. In: M.A. Rankin (editor), Migrations: Mechanisms and Adaptive Significance. Contributions in Marine Science, Supplement to Vol. 27: 353-366.

Quinn, T.P. Salmon homing: Is the puzzle complete? Review of A.D. Hasler and A.T. Scholz, Olfactory Imprinting and Homing in Salmon. Environmental Biology of Fishes 12: 315-317.

Quinn, T.P. and C.A. Busack. Chemosensory recognition of siblings in juvenile coho salmon (Oncorhynchus kisutch). Animal Behaviour 33: 51-56.

Brannon, E.L., R.P. Whitman and T.P. Quinn. Responses of returning adult coho salmon to home water and population-specific odors. Transactions of the American Fisheries Society 113: 374-377.

Ogden, J.C. and T.P. Quinn. Migration in coral reef fishes: Ecological significance and orientation mechanisms. In: J.D. McCleave,G.P. Arnold, J.J. Dodson and W.H. Neill (editors), Mechanisms of Migration in Fishes. Plenum Press, New York, pp. 293-308.

Quinn, T.P. An experimental approach to fish compass and map orientation. In: J.D. McCleave, G.P. Arnold, J.J. Dodson and W.H. Neill (editors), Mechanisms of Migration in Fishes. Plenum Press, New York, pp. 113-123.

Quinn, T.P. Homing and straying in Pacific salmon. In: J.D. McCleave, G.P. Arnold, J.J. Dodson and W.H. Neill (editors), Mechanisms of Migration in Fishes. Plenum Press, New York, pp. 357-362.

Quinn, T.P. and K. Fresh. Homing and straying in chinook salmon (Oncorhynchus tshawytscha) from Cowlitz River Hatchery, Washington. Canadian Journal of Fisheries and Aquatic Sciences 41: 1078-1082.

Quinn, T.P. and C. Groot. Pacific salmon (Oncorhynchus) migrations: Orientation vs. random movement. Canadian Journal of Fisheries and Aquatic Sciences 41: 1319-1324.

Quinn, T.P. and C. Groot. The effect of water flow rate on bimodal orientation of juvenile chum salmon, Oncorhynchus keta. Animal Behaviour 32: 628-629.

Quinn, T.P. and J.C. Ogden. Field evidence of compass orientation in migrating juvenile grunts (Haemulidae). Journal of Experimental Marine Biology and Ecology 81: 181-192.

Quinn, T.P., E.L. Brannon and R.P. Whitman. Pheromones and the water source preferences of adult coho salmon (Oncorhynchus kisutch). Journal of Fish Biology 22: 677-684.

Quinn, T.P. and C. Groot. Orientation of chum salmon (Oncorhynchus keta) after internal and external magnetic field alteration. Canadian Journal of Fisheries and Aquatic Sciences 40: 1598-1606.

Quinn, T.P. A model for salmon navigation on the high seas. In: E.L. Brannon and E.O. Salo (editors), Salmon and Trout Migratory Behavior Symposium. School of Fisheries, University of Washington, pp. 229-237.

Quinn, T.P. Intra-specific differences in sockeye salmon fry compass orientation mechanisms. In: E.L. Brannon and E.O. Salo (editors), Salmon and Trout Migratory Behavior Symposium. School of Fisheries, University of Washington, pp. 79-85.

Quinn, T.P. and E.L. Brannon. The use of celestial and magnetic cues by orienting sockeye salmon smolts. Journal of Comparative Physiology 147: 547-552.

Whitman, R.P., T.P. Quinn and E.L. Brannon. The influence of suspended volcanic ash on the homing behavior of adult chinook salmon. Transactions of the American Fisheries Society 111: 63-69.

Brannon, E.L., T.P. Quinn, G.L. Lucchetti and B.D. Ross. Compass orientation of sockeye salmon from a complex river system. Canadian Journal of Zoology 59: 1548-1553.

Quinn, T.P. Compass orientation of juvenile sockeye salmon (Oncorhynchus nerka). Doctoral dissertation, University of Washington.

Quinn, T.P., R.T. Merrill and E.L. Brannon. Magnetic field detection in sockeye salmon. Journal of Experimental Zoology 217: 137-142.

Quinn, T.P. Evidence for celestial and magnetic compass orientation in lake migrating sockeye salmon fry. Journal of Comparative Physiology 137: 243-248.

Quinn, T.P. Locomotor responses of juvenile blind cave fish, Astyanax jordani, to the odors of conspecifics. Behavioral and Neural Biology 29: 123-127.

Quinn, T.P., B.S. Miller and R.C. Wingert. Depth distribution and seasonal and diel movements of ratfish, Hydrolagus colliei, in Puget Sound, Washington. Fishery Bulletin 78: 816-821.

Quinn, T.P. The role of the odors of conspecifics in the locomotor behavior of the blind cave characin, Astyanax jordani. Master’s thesis, University of Washington.

Quinn, T., G.W. Esch, T.C. Hazen and J.W. Gibbons. Long range movement and homing by largemouth bass (Micropterus salmoides) in a thermally altered reservoir. Copeia 1978: 542-545.

 

The Behavior and Ecology of Pacific Salmon and Trout: 2018 Revised Edition

A completely revised edition of the 2005 book, “The Behavior and Ecology of Pacific Salmon and Trout” is available through UW Press.

This new edition features many new, outstanding photographs, updated information, and enhanced coverage of trout and char as well as Pacific salmon. There have been many interesting and important discoveries since the first edition, and the new edition presents them. The book is currently available in paperback, with an e-book option also available.

The Behavior and Ecology of Pacific Salmon and Trout should be on the shelf of anyone with a professional or personal interest in Pacific salmon and trout. Written in a technically accurate but engaging style, it appeals to a wide range of readers, including students, anglers, biologists, conservationists, legislators, and armchair naturalists.

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Courses

Salmonid Behavior and Life History – FISH 450

Goals

 

The purpose of this class is to familiarize students with the life history, behavior, and ecology of Pacific salmon and trout. The lectures provide detailed information on the marine distribution, homing migration, energetics, and spawning behavior of adult salmon, the ecological processes linked to their carcasses, the incubation and emergence of their offspring from gravel nests, the ecology of juveniles in streams and lakes, their seaward migration, and ecology at sea.

This class is fast-paced and information rich, and it is expected that students are already familiar with fish biology and ecological principles, and have also had experience processing data and writing scientific papers.

  • Introduction, goals and course schedule
  • General life history & distribution of Oncorhynchus, Salmo and Salvelinus
  • Life histories and identification, continued
  • Life histories compared with other N. A. fishes
  • Global patterns of anadromy and catadromy; Speciation, glaciation
  • Oceanic migration and orientation
  • Coastal and estuarine migrations
  • Upriver migrations, energetics, hormones
  • Homing – evidence and patterns
  • Homing mechanisms
  • Selection of spawning site and female reproductive behavior
  • Reproductive success of males
  • Ecology of adult salmon: predators, scavengers and nutrient cycling
  • Embryo mortality and incubation schedules
  • Emergence and initial fry movements
  • Sockeye fry in lakes: predation and feeding
  • Salmon and trout in streams Part I & II
  • Smolt transformation & physiology
  • Seaward migration: timing, orientation
  • Estuarine residence and migration
  • Early marine migrations and distribution
  • Survival at sea
  • Feeding and growth at sea
  • Age and size at maturity
  • Population structure and local adaptation
  • Successes and failures of transplants
  • Effects of land-use activities on salmonids
  • Wild salmon and hatcheries
  • The future of salmon in a changing world

Aquatic Ecological Research in Alaska – FISH 491

Background

FISH 491 students in Bristol Bay, Alaska

The University of Washington and the School of Aquatic & Fishery Sciences (SAFS), in particular, are interested in closely connecting undergraduate teaching and research.

As one step towards this goal, the school offers a course in aquatic ecology, started in summer 1999, at the Fisheries Research Institute’s field stations in Bristol Bay, Alaska

Goals

The goal of the course is to provide a small number of students with direct, hands-on training in the theories and techniques of research in aquatic ecology. We make use of the well-equipped field camps and diverse aquatic environments to give students formal course material (i.e., lectures and instruction manuals), directed research and independent projects.

Course Structure

Fish 491 students in Bristol Bay, Alaska

FISH 491 students in Bristol Bay, Alaska

All students will enroll in Fish 491 for 4 credits (graded) in summer quarter and will jointly participate in field and laboratory research at lakes Aleknagik and Iliamna. The course will run for about about a month, starting in mid-July. During that time students will receive instruction in limnology, juvenile fish ecology, spawning behavior and life history of adult salmon, population dynamics and fishery management. Students will have access to the long-term data sets collected at the camps, and will write papers on limnology, fish behavior and population dynamics.

  • Mariah Meek: Temperature, ice coverage, and the growth of juvenile Iliamna Lake sockeye salmon
  • Kristi Overberg: Comparison of adult sockeye salmon morphology among rivers
  • Erin Meehan: Scavenging of sockeye salmon carcasses by blowfly maggots
  • Donna Hauser: Haul out patterns and diet of harbor seals in Iliamna Lake
  • Ranae Holland: Patterns of bear activity on Hansen Creek
  • Sue Johnson: Armored for battle: teeth and skin of sockeye salmon populations
  • Harry Rich, Jr.: Longevity, mortality, and movement of jacks on the spawning grounds.
  • Michael Cotter: Diet overlap in juvenile Dolly Varden and coho salmon in streams
  • Casey Ruff: Diet overlap and prey selectivity of sockeye salmon and 3 spine sticklebacks
  • Erin Lowery: Meristic and morphometric comparison of Arctic charr and Dolly Varden
  • Jennifer Marsh: Diel vertical migration of zooplankton in Iliamna Lake
  • Daniel Okamoto: Community composition of resident stream fishes
  • Chris Allen: Alternate reproductive tactics and large and jack sockeye salmon
  • Anna Buettner: Prevalence and infestation of the parasite Schistocephalus in sticklebacks
  • Tanya Kleisborg: Egg retention and prespawning mortality of female sockeye salmon.
  • Katy Shipe: Effects salmon spawning on the community of stream insects
  • Jamie Thompson: Abundance and ontogenetic habitat segregation in Dolly Varden char
  • Eahn Holm: egg size/fecundity tradeoff as a function of age and size in sockeye salmon
  • Trevor Hutton: scavenging of sockeye salmon by bears
  • Andrea Odle: diet of sculpins on Iliamna Lake beaches
  • Kristin Omori: relationship between coho salmon fry size and ability to eat sockeye eggs
  • Leslie Jensen: growth of freshwater mussels in relation to climate variation among years
  • Gretchen Theusen: food habits and growth of northern pike
  • Hannah Barrett: Attraction of salmonids to the chemical traces of salmon eggs
  • Brittany Cummings: Effects of spawning salmon on the size of emerging aquatic insects
  • Ke’ale Louie: Energy content of different tissues of sockeye salmon
  • Jeanelle Miller: Effect of hyporheic flow on salmon spawning distribution Hansen Creek
  • Brian Harmon: Effects of cestode parasites on sculpin age, growth, and survival
  • Rosalind Huang: Activity patterns and feeding of Arctic terns at Lake Aleknagik
  • Jenna Keeton: Color change and substrate color preference by sculpins
  • Brendan Smith: Transportation of salmon carcasses into the forest by bears
  • Susan Harris: Spatial distribution of salmon density and bear predation on Hansen Creek
  • Nirupam Nigam: Behavior of gulls at the mouth of Hansen Creek

Instructors

The primary instructors will be Dr. Thomas Quinn (SAFS), Dr. Daniel Schindler (SAFS) and Dr. Ray Hilborn (SAFS). There will also be a teaching assistant to help students with the fieldwork and their independent projects.

FISH 491 students boarding a float plane

Undergraduate Endowment

Thomas Quinn and his wife, Sandra O’Neill have set up an endowment to enable undergraduate students to get out of the classroom and into the field, and have the kind of transformative experience that only field work can provide. Students are eager for the hands-on field work that provides a lifetime of memories and helps define career pathways.

If you share a love of field research and student learning, and wish to help build the next generation of scientists, teachers, and informed citizens, please consider contributing to this fund. Contact Tom Quinn if you have any questions, and use the below to start the process of contributing to this fund. Any amount will help.

Contribute to the next generation of scientists via the O’Neill Quinn Aquatic and Fishery Sciences Endowed Immersive Learning Fund